Английская Википедия:2012 in archosaur paleontology

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Шаблон:Year nav topic20Шаблон:Portal The year 2012 in Archosaur paleontology was eventful. Archosaurs include the only living dinosaur group — birds — and the reptile crocodilians, plus all extinct dinosaurs, extinct crocodilian relatives, and pterosaurs. Archosaur palaeontology is the scientific study of those animals, especially as they existed before the Holocene Epoch began about 11,700 years ago. The year 2012 in paleontology included various significant developments regarding archosaurs.

This article records new taxa of fossil archosaurs of every kind that have been described during the year 2012, as well as other significant discoveries and events related to paleontology of archosaurs that occurred in the year 2012.

Pseudosuchians

Research

• The postcranial skeleton of Sebecus icaeorhinus is described by Diego Pol et al. (2012).
• Skull anatomy of Dakosaurus and Plesiosuchus is described by Mark T. Young et al. (2012).
• Kemkemia auditorei, previously thought to be a theropod dinosaur, is reinterpreted as a member of Crocodyliformes by Gabriel Lio et al. (2012).
• A study including a supertree of Crocodyliformes is published by Mario Bronzati, Felipe Chinaglia Montefeltro and Max C. Langer (2012).
• Skull remains of a member of the genus Gavialis belonging or related to the species Gavialis bengawanicus are described from the Early Pleistocene of Thailand by Martin et al. (2012).

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Aegisuchus	Gen. et sp. nov	Valid	Holliday & Gardner	Cenomanian	Kem Kem Formation		An aegyptosuchid eusuchian.	Файл:Aegisuchus.png Aegisuchus witmeri.
Aetobarbakinoides	Gen. et sp. nov	Valid	Desojo, Ezcurra & Kischlat	Late Carnian or early Norian	Santa Maria Formation		An aetosaur.
Apachesuchus	Gen. et sp. nov	Valid	Spielmann & Lucas	Late Triassic			An aetosaur. The type species is Apachesuchus heckerti.
Barreirosuchus	Gen. et sp. nov	Valid	Iori & Garcia	Turonian to Santonian	Adamantina Formation		A trematochampsid. The type species is Barreirosuchus franciscoi.
Borealosuchus threeensis	Sp. nov	Valid	Brochu, Parris, Grandstaff, Denton & Gallagher	Maastrichtian or Danian			A crocodilian from New Jersey.
Bystrowisuchus	Gen. et sp. nov	Valid	Sennikov	Early Triassic			A ctenosauriscid. The type species is Bystrowisuchus flerovi.
Crocodylus thorbjarnarsoni	sp nov	Valid	Brochu & Storrs	Pliocene to Pleistocene			A species of Crocodylus.	Файл:Crocodylus thorbjarnarsoni.jpg Crocodylus thorbjarnarsoni.
Gasparinisuchus	Gen. et sp. nov	Valid	Martinelli et al.	Late Cretaceous			A peirosaurid. The type species is Gasparinisuchus peirosauroides.

Newly named basal dinosauriforms

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Diodorus	Gen. et sp. nov	Valid	Kammerer, Nesbitt, & Shubin	?Carnian-Norian	Timezgadiouine Formation		A silesaurid.

Non-avian dinosaurs

Research

• A study including a large phylogenetic analysis of non-coelurosaurian tetanuran theropod dinosaurs is published by Matthew T. Carrano, Roger B. J. Benson and Scott D. Sampson (2012).
• A study including a systematic revision of the family Dromaeosauridae and a large phylogenetic analysis of paravian theropod dinosaurs (both avian and non-avian) is published by Alan Turner, Peter Makovicky and Mark Norell (2012).
• A specimen of Microraptor is described by Quanguo Li et al. (2012), with melanosome imprints making it possible to determine the plumage coloration and iridescence in the plumage.
• A new specimen of Austroraptor cabazai is described by Philip J. Currie and Ariana Paulina Carabajal (2012).
• A specimen of Mei long is described by Chunling Gao et al. (2012).
• A study of Khaan mckennai was published by Amy M. Balanoff and Mark Norell (2012).
• A new information on Yixianosaurus longimanus is published by T. Alexander Dececchi, Hans C. E. Larsson and David W. E. Hone (2012).
• Three specimens of Ornithomimus with evidence of feathers are described by Darla K. Zelenitsky et al. (2012).
• A new information on the anatomy of the holotype specimen of Nqwebasaurus thwazi is published by Jonah N. Choiniere, Catherine A. Forster and William J. de Klerk (2012).
• A study of Alioramus was published by Stephen Brusatte, Thomas D. Carr and Mark Norell (2012).
• Abdominal contents of two specimens of Sinocalliopteryx gigas are described by Lida Xing et al. (2012).
• A study of musculoskeletal anatomy, three-dimensional body proportions and body mass evolution in allosauroid theropod dinosaurs is published by Karl T. Bates, Roger B. J. Benson, and Peter L. Falkingham  (2012).
• A study of the braincase of Sinraptor dongi is published by Ariana Paulina Carabajal and Philip J. Currie (2012).
• A study of Kelmayisaurus petrolicus, interpreting it as likely to be a member of Carcharodontosauridae, is published by Stephen L. Brusatte, Roger B.J. Benson and Xing Xu (2012).
• Description of pectoral girdle and forelimb of Majungasaurus crenatissimus is published by Sara H. Burch and Matthew T. Carrano (2012).
• A study of Early Cretaceous Australian theropod dinosaurs was published by Roger B. J. Benson et al. (2012).
• A study including a phylogenetic analysis of titanosauriform sauropod dinosaurs is published by Michael D'Emic (2012).
• The postcranial skeletal pneumaticity in the skeletons of five taxa of early sauropodomorph dinosaurs is described by Adam M. Yates, Mathew J. Wedel and Matthew F. Bonnan (2012).
• The study on the presumed course of the recurrent laryngeal nerve in sauropod dinosaurs is published by Mathew J. Wedel (2012).
• Pachysuchus, previously thought to be an Early Jurassic phytosaur, is reinterpreted as a sauropodomorph dinosaur by Paul M. Barrett and Xu Xing (2012).
• A study of vertebral laminae of sauropod dinosaurs is published by Jeffrey A. Wilson (2012).
• A study on the neural spine bifurcation in diplodocoid sauropod dinosaurs is published by D. Cary Woodruff and Denver W. Fowler (2012).
• A study of the postcranial skeletal pneumaticity in the skeletons of Saltasaurus, Neuquensaurus and Rocasaurus is published by Ignacio A. Cerda, Leonardo Salgado and Jaime E. Powell  (2012).
• A study of Early Cretaceous sauropod dinosaurs from North America is published by Michael D. D’Emic and Brady Z. Foreman (2012). Among other findings, additional sauropod material from the Cloverly Formation of Wyoming was referred to Sauroposeidon, Paluxysaurus was synonymized with Sauroposeidon, Rugocaudia was considered a nomen dubium and the cause of the North American sauropod extinction in the middle of the Cretaceous period was discussed.
• The first sauropod dinosaur (a member of Titanosauria) from Antarctica described by Ignacio A. Cerda et al.  (2012).
• A study of biomechanics, pectoral girdle articulation and body mass of the Triassic dinosaurs from Brazil (Staurikosaurus, Saturnalia, Pampadromaeus, Guaibasaurus and Unaysaurus) is published by Rafael Delcourt, Sergio A. K. de Azevedo, Orlando N. Grillo and Fernanda O. Deantoni (2012).
• Anatomy of Fruitadens haagarorum is described by Richard J. Butler et al. (2012).
• A study including a phylogenetic analysis of ankylosaurian dinosaurs is published by Richard S. Thompson et al. (2012).
• A study including a phylogenetic analysis of iguanodontian ornithopod dinosaurs is published by Andrew T. McDonald (2012).
• A study of the forearm orientation in hadrosaurids is published by Phil Senter (2012).
• Skin impressions of two different species of Saurolophus are described by Phil Bell (2012).
• A new description of Eolambia caroljonesa is published by Andrew T. McDonald et al. (2012).
• A study of anatomy and relationships of Bolong yixianensis is published by Wu Wenhao and Pascal Godefroit (2012).
• An overview of Early Cretaceous iguanodontian ornithopod dinosaurs of England and Belgium is published by David B. Norman (2012).
• A study of the bone histology of Tenontosaurus tilletti is published by Sarah Werning (2012).
• A study of the bone histology of Dysalotosaurus lettowvorbecki is published by Tom R. Hübner (2012).
• A study questioning the interpretation of Torosaurus as a junior synonym and a growth stage of Triceratops was published by Nicholas R. Longrich and Daniel J. Field (2012).
• A study of some of the earliest known dinosaur assemblages is published by Martín D. Ezcurra (2012). Among other findings, Teyuwasu is interpreted as a member of the clade Dinosauriformes of uncertain phylogenetic placement and a nomen dubium.
• A study of anatomical variability exhibited by major dinosaur groups living during the latest Cretaceous is published by Stephen L. Brusatte et al. (2012).
• A study of biodiversity of late Maastrichtian non-avian dinosaurs is published by Jean Le Loeuff (2012).

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Alnashetri	Gen. et sp. nov	Valid	Makovicky, Apesteguía & Gianechini	Late Cretaceous (Cenomanian or Turonian)	Candeleros Formation		A coelurosaurian, a probable relative of alvarezsaurids. The type species is Alnashetri cerropoliciensis.
Astrophocaudia	Gen. et sp. nov	Valid	D’Emic	Early Cretaceous	Trinity Group	Шаблон:Flag (Шаблон:Flag)	A titanosauriform sauropod, a member of the clade Somphospondyli. The type species is Astrophocaudia slaughteri.
Batyrosaurus	Gen. et sp. nov	Valid	Godefroit et al.	Late Cretaceous			A basal hadrosauroid. The type species is Batyrosaurus rozhdestvenskyi.
Bicentenaria	Gen. et sp. nov	Valid	Novas et al.	Cenomanian	Candeleros Formation		A coelurosaur. The type species is Bicentenaria argentina.
Bonapartenykus	Gen. et sp. nov	Valid	Agnolin, Powell, Novas, & Kundrát	Campanian/Maastrichtian	Allen Formation		An alvarezsaurid.
Comahuesaurus	Gen. et sp. nov	Valid	Carballido et al.	Early Cretaceous (Aptian or Albian)	Lohan Cura Formation		A rebbachisaurid sauropod. The type species is Comahuesaurus windhauseni.
Coronosaurus	Gen. et comb. nov	Valid	Ryan, Evans & Shepherd	Late Cretaceous	Oldman Formation		A centrosaurine ceratopsian, a new genus for "Centrosaurus" brinkmani Ryan & Russell (2005).	Файл:Coronosaurus NT small.jpg Coronosaurus
Darwinsaurus	Gen. et sp. nov	Valid	Paul	Early Cretaceous (early Valanginian)	Wadhurst Clay Formation		A styracosternan ornithopod. The type species is Darwinsaurus evolutionis, and the holotype is an associated skeleton that includes material catalogued under numbers NHMUK R1831 (a dentary), R1833, R1835 and R1836; this skeleton is assigned by Norman (2010) to Hypselospinus fittoni.
Elaltitan	Gen. et sp. nov	Valid	Mannion & Otero	Middle Cenomanian to Turonian	Bajo Barreal Formation		A lithostrotian titanosaur. The type species is Elaltitan lilloi.
Eoabelisaurus	Gen. et sp. nov	Valid	Pol & Rauhut	Aalenian or Bajocian	Cañadon Asfalto Formation		An abelisaurid. The type species is Eoabelisaurus mefi.	Файл:Eoabelisaurus restoration.png Eoabelisaurus
Gideonmantellia	Gen. et sp. nov	Valid	Ruiz-Omeñaca et al.	Barremian	Camarillas Formation		A basal ornithopod. The type species is Gideonmantellia amosanjuanae.	Файл:Gideonmantellia.jpg Gideonmantellia
Gryphoceratops	Gen. et sp. nov	Valid	Ryan, Evans, Currie, Brown, & Brinkman	Late Santonian	Milk River Formation		A leptoceratopsid ceratopsian.
Hexing	Gen. et sp. nov	Valid	Jin, Chen & Godefroit	Early Valanginian to early Barremian	Yixian Formation		A basal ornithomimosaur. The type species is Hexing qingyi.
Huehuecanauhtlus	Gen. et sp.	Valid	Ramírez-Velasco et al.	Santonian			A basal hadrosauroid. The type species is Huehuecanauhtlus tiquichensis.
Huxleysaurus	Gen. et com. nov	Valid	Paul	Early Cretaceous (early Valanginian)	Wadhurst Clay Formation		A styracosternan ornithopod, a new genus for "Iguanodon" hollingtoniensis Lydekker (1889); this species is considered by Norman (2010) to be a junior synonym of Hypselospinus fittoni.
Ichthyovenator	Gen. et sp. nov	Valid	Allain et al.	Late Early Cretaceous	Savannakhet Basin		A spinosaurid. The type species is Ichthyovenator laosensis.
Kaatedocus	Gen. et sp. nov	Valid	Tschopp & Mateus	Late Jurassic (Kimmeridgian)	Morrison Formation		A diplodocine diplodocid sauropod. The type species is Kaatedocus siberi.	Файл:Kaatedocus - Muse front.jpg Kaatedocus
Kundurosaurus	Gen. et sp. nov	Valid	Godefroit, Bolotsky & Lauters	Late Cretaceous, possibly late Maastrichtian	Udurchukan Formation		A saurolophine hadrosaurid. The type species is Kundurosaurus nagornyi.	Файл:Kundurosaurus.tif Kundurosaurus.
Lapampasaurus	Gen. et sp. nov	Valid	Coria, Riga & Casadío	Late Cretaceous (late Campanian or early Maastrichthian)	Allen Formation		A hadrosaurid. The type species is Lapampasaurus cholinoi.
Latirhinus	Gen. et sp. nov	Valid	Prieto-Márquez & Brañas	Late Cretaceous (late Campanian)	Cerro del Pueblo Formation		A hadrosaurid of uncertain phylogenetic placement; originally described as a saurolophine, but subsequently reinterpreted as a lambeosaurine. The type species is Latirhinus uitstlani.
Magnapaulia	Gen. et comb. nov	Valid	Prieto-Márquez, Chiappe & Joshi	Late Campanian	El Gallo Formation		A lambeosaurine hadrosaurid, a new genus for "Lambeosaurus" laticaudus (Morris, 1981).	Файл:Magnapaulia.tif Magnapaulia.
Mantellodon	Gen. et sp. nov	Valid	Paul	Early Cretaceous (early Aptian)	Lower Greensand Formation		A styracosternan ornithopod. The type species is Mantellodon carpenteri and the holotype is the specimen NHMUK R3741 consisting of partial associated skeleton (also known as Gideon Mantell's "Mantel-piece")
Martharaptor	Gen. et sp. nov	Valid	Senter, Kirkland & DeBlieux	Early Aptian	Cedar Mountain Formation		A possible therizinosauroid. The type species is Martharaptor greenriverensis.	Файл:Martharaptor greenriverensis.png Martharaptor.
Microraptor hanqingi	sp nov	Valid	Gong et al.	Early Cretaceous			A species of Microraptor. Its status as a distinct species is disputed, with Pei et al. (2014) considering the holotype specimen's morphology not to be distinguishable from other Microraptor specimens.
Mochlodon vorosi	Sp. nov	Valid	Ősi et al.	Santonian	Csehbánya Formation		A rhabdodontid ornithopod, a species of Mochlodon.
Ningyuansaurus	Gen. et sp. nov	Valid	Ji et al.	Early Cretaceous	Yixian Formation		An oviraptorosaur. The type species is Ningyuansaurus wangi.
Ostafrikasaurus	Gen. et sp. nov	Valid	Buffetaut	Late Jurassic	Tendaguru Formation		A spinosaurid. The type species is Ostafrikasaurus crassiserratus.
Pachyrhinosaurus perotorum	sp nov	Valid	Fiorillo & Tykoski	Maastrichtian	Prince Creek Formation		Third known species of Pachyrhinosaurus.
Pegomastax	Gen. et sp. nov	Valid	Sereno	Hettangian to Sinemurian	Elliot Formation		A heterodontosaurid. The type species is Pegomastax africana.	Файл:Pegomastax dino.jpg Pegomatax
Philovenator	Gen. et sp. nov	Valid	Xu et al.	Campanian	Wulansuhai Formation		A troodontid. Its type species is Philovenator curriei.	Файл:Philovenator curriei life restoration..png Philovenator
Proa	Gen. et sp. nov	Valid	McDonald et al.	Early Cretaceous (early Albian)	Escucha Formation		An iguanodont ornithopod closely related to Iguanodon and the clade Hadrosauroidea. The type species is Proa valdearinnoensis.
Rugocaudia	Gen. et sp. nov	Valid	Woodruff	Aptian or Albian	Cloverly Formation		A titanosauriform sauropod. The type species is Rugocaudia cooneyi.
Sauroniops	Gen. et sp. nov	Valid	Cau, Dalla Vecchia & Fabbri	Cenomanian	Kem Kem Beds		A carcharodontosaurid. The type species is Sauroniops pachytholus. Announced in 2012; the final version of the article naming it was published in 2013.
Sciurumimus	Gen. et sp. nov	Valid	Rauhut et al.	Upper Kimmeridgian			A megalosauroid. The type species is Sciurumimus albersdoerferi.	Файл:Xaveropterus 2.jpg Sciurumimus.
Unescoceratops	Gen. et sp. nov	Valid	Ryan, Evans, Currie, Brown, & Brinkman	Late Campanian	Dinosaur Park Formation		A leptoceratopsid ceratopsian.
Xenoceratops	Gen. et sp. nov	Valid	Ryan, Evans & Shepherd	Late Cretaceous (middle Campanian)	Foremost Formation		A centrosaurine ceratopsian. The type species is Xenoceratops foremostensis.	Файл:Xenoceratops NT small.jpg Xenoceratops
Yueosaurus	Gen. et sp. nov	Valid	Zheng, Jin, Shibata, Azuma, & Yu	Aptian-Cenomanian	Liangtoutang Formation		A basal ornithopod.
Yurgovuchia	Gen. et sp. nov	Valid	Senter et al.	Early Cretaceous, possibly Barremian	Cedar Mountain Formation		A dromaeosaurid. The type species is Yurgovuchia doellingi.	Файл:Reconstruction of Yurgovuchia doellingi.png Yurgovuchia.
Yutyrannus	Gen. et sp. nov	Valid	Xu et al.	Early Cretaceous	Yixian Formation		A basal tyrannosauroid. The type species is Yutyrannus huali.	Файл:Feathered tyrant by pilsator-d4vbemk.jpg Yutyrannus.

Newly named birds

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Aegolius gradyi	Sp. nov.	Valid	Storrs L. Olson	Pleistocene to Holocene			A Strigidae.
Aegypius jinniushanensis	Sp. nov.	Valid	Zhang Zihui Huang Yunping Helen F. James Hou Lianhai	Middle Pleistocene			An Accipitridae.
Aix praeclara	Sp. nov.	Valid	Nikita V. Zelenkov Evgeny N. Kurochkin	Middle Miocene	Oshin Formation		An Anatidae.
Alauda xerarvensis	Sp. nov.	Valid	Zlatozar Boev	Late Pliocene	Middle Villafranchian		An Alaudidae.
Anchigyps voorhiesi	Gen. nov. et Sp. nov.	Valid	Zhang Zihui Alan Feduccia Helen F. James	Late Miocene. late Clarendonian	Ash Hollow Formation	Шаблон:Flag:	An Accipitridae close in morphology to the Old World Vultures. The type species of the new genus.
Anhinga walterbolesi	Sp. nov.	Valid	Trevor H. Worthy	Late Oligocene or Early Miocene	Etadunna Formation, 24-26 Ma	Шаблон:Flag:	An Anhingidae.
Anthus antecedens	Sp. nov.	Valid	Jenö Kessler János Hír	Middle Miocene	MN 7-8		A Motacillidae.
Athene vallgornerensis	Sp. nov.	Valid	Carmen Guerra Pere Bover Josep A. Alcover	Early Pleistocene	Cave deposits	Шаблон:Flag:	A Strigidae.
Australopicus nelsonmandelai	Gen. nov. et Sp. nov.	Valid	Albrecht Manegold Antoine Louchart	Early Pliocene	Varswater Formation		A Picidae, this is the type species of the new genus.
Avolatavis tenens	Gen. nov. et Sp. nov.	Valid	Daniel T. Ksepka Julia A. Clarke	Eocene	Green River Formation	Шаблон:Flag:	A stem parrot, a member of Pan-Psittaciformes. The type species of the new genus.
Becassius charadriioides	Gen. nov. et Sp. nov.	Valid	Vanesa L. De Pietri Gerald Mayr	Early Miocene	Saint-Gérard-le-Puy, MN 2a		Originally described as a member of the family Scolopacidae or a relative of the Scolopacidae; De Pietri, Mayr & Scofield (2019) transferred this species to the family Glareolidae. The type species of the new genus.
Bombycilla hamori	Sp. nov.	Valid	Jenö Kessler János Hír	Early Miocene	MN 5		A Bombycillidae.
Brodavis americanus	Gen. nov. et Sp. nov.	Valid	Larry D. Martin Evgeny N. Kurochkin Tim T. Tokaryk	Late Cretaceous,	Maastrichtian Frenchman Formation	Шаблон:Flag:	A Hesperornithiformes Fürbringer, 1888, Brodavidae Martin, Kurochkin et Tokaryk, 2012, this is the type species of the new genus.
Brodavis baileyi	Sp. nov.	Valid	Larry D. Martin Evgeny N. Kurochkin Tim T. Tokaryk	Late Cretaceous,	Maastrichtian Hell Creek Formation	Шаблон:Flag:	A Hesperornithiformes Fürbringer, 1888, Brodavidae Martin, Kurochkin et Tokaryk, 2012.
Brodavis mongoliensis	Sp. nov.	Valid	Larry D. Martin Evgeny N. Kurochkin Tim T. Tokaryk	Late Cretaceous,	Maastrichtian Nemegt Formation		A Hesperornithiformes Fürbringer, 1888, Brodavidae Martin, Kurochkin et Tokaryk, 2012.
Certhia janossyi	Sp. nov.	Valid	Jenö Kessler János Hír	Late Miocene	MN 9		Originally described as a species of Certhia; Zelenkov (2017) transferred this species to a new genus Sylvosimadaravis within the superfamily Sylvioidea.
Chenoanas deserta	Gen. nov. et Sp. nov.	Valid	Nikita V. Zelenkov	Middle Miocene	Oshin Formation		An Anatidae, this is the type species of the new genus.
Cinclus major	Sp. nov.	Valid	Jenö Kessler János Hír	Early Miocene	MN 5		A Cinclidae.
Circaetus rhodopensis	Sp. nov.	Valid	Zlatozar Boev	Late Miocene	Tulorian = Late Meotian, Late MN 11, ca. 7,5 Mya		An Accipitridae.
Crypturellus reai	Sp. nov.	Valid	Robert M. Chandler	Miocene (Santacrucian)	Santa Cruz Formation		A Tinamidae.
?Elorius limosoides	Sp. nov.	Valid	Vanessa L. De Pietri Gerald Mayr	Early Miocene	Saint-Gérard-le-Puy, MN 2a		A Scolopacidae, possibly a species of Elorius.
Emberiza bartkoi	Sp. nov.	Valid	Jenö Kessler János Hír	Early Miocene	MN 5		An Emberizidae.
Eremarida xerophila	Gen. nov. et Sp. nov.	Valid	Zlatozar Boev	Late Miocene	Gnilyane Formation		An Alaudidae, this is the type species of the new genus.
Eremophila prealpestris	Sp. nov.	Valid	Zlatozar Boev	Late Pliocene	Middle Villafranchian, MP 17, 2,23-2,40 Mya		An Alaudidae.
Erithacus horusitzkyi	Sp. nov.	Valid	Jenö Kessler János Hír	Middle Miocene	MN 7/8		A Muscicapidae, an Erithacus robin.
Galerida bulgarica	Sp. nov.	Valid	Zlatozar Boev	Late Pliocene	Middle Villafranchian		An Alaudidae.
Galerida cserhatensis	Sp. nov.	Valid	Jenö Kessler János Hír	Early Miocene	MN 5		An Alaudidae.
Heteroanser vicinus	Gen. nov. et Comb. nov.	Valid	Nikita V. Zelenkov	Late Miocene	Hyargas Nuur Formation		An Anatidae a new genus for "Heterochen" vicinus Kurochkin, 1976.
Hirundo selengenica	Sp. nov.	Valid	Nikita V. Zelenkov Evgeny N. Kurochkin	Late Pliocene	Chikoi Formation		A Hirundinidae.
Jeholornis palmapenis	Sp. nov.	Valid	Jingmai K. O'Connor Sun Chengkai Xu Xing Wang Xiaolin Zhou Zhonghe	Early Aptian	Jiufotang Formation		A Jeholornithidae Zhou et Zhang, 2006.
Kairuku grebneffi	Sp. nov.	Valid	Daniel T. Ksepka R. Ewan Fordyce Tatsuro Ando Craig M. Jones	Oligocene	Dutroonian		A Spheniscidae.
Kairuku waitaki	Gen. nov. et Sp. nov	Valid	Daniel T. Ksepka R. Ewan Fordyce Tatsuro Ando Craig M. Jones	Oligocene	Late Whaingaroan or Early Duntroonian		A Spheniscidae, the type species of the genus.
Lanius schreteri	Sp. nov.	Valid	Jenö Kessler János Hír	Middle Miocene	MN 7-8		A Laniidae.
Leptoptilos lüi	Sp. nov.	Not valid	Zhang Zihui Huang Yunping Helen F. James Hou Lianhai	Middle Pleistocene	Cave deposits		A Ciconiidae, the name is not valid because of the "ü" in it, that must, of course be a plain "u".
Lullula balcanica	Sp. nov.	Valid	Zlatozar Boev	Late Pliocene	Middle Villafranchian		An Alaudidae.
Lullula neogradensis	Sp. nov.	Valid	Jenö Kessler János Hír	Middle Miocene	MN 7/8		An Alaudidae.
Lullula slivnicensis	Sp. nov.	Valid	Zlatozar Boev	Late Pliocene	Early Late Villafranchian, MN 18		An Alaudidae, a relative of the woodlark.
Luscinia praeluscinia	Sp. nov.	Valid	Jenö Kessler János Hír	Early Miocene	MN 5		A Muscicapidae.
Lutetodontopteryx tethyensis	Gen. nov. et Sp. nov.	Valid	Gerald Mayr Evgenij Zvonok	Middle Eocene	Early Lutetian		A Pelagornithidae, this is the type species of the new genus.
Melanocorypha donchevi	Sp. nov.	Valid	Zlatozar Boev	Late Pliocene	Middle Villafranchian		An Alaudidae.
Melanocorypha serdicensis	Sp. nov.	Valid	Zlatozar Boev	Late Miocene	Gnilyane Formation, Middle Late Pontian		An Alaudidae.
Mioquerquedula minutissima	Gen. nov. et Sp. nov.	Valid	Nikita V. Zelenkov Evgeny N. Kurochkin	Middle Miocene, possibly also Late Miocene	Gobi-Altai Aimag, Shargyn Govi, Sharga location	Шаблон:Flag	An Anatidae, this is the type species of the new genus, the new genus also contains "Anas" velox Milne-Edwards, 1868 as a Comb. nov. from France, Germany, Romania and ?Hungary.
Muscicapa leganyii	Sp. nov.	Valid	Jenö Kessler János Hír	Middle Miocene	MN 7/8		A Muscicapidae.
Otus mauli	Sp. nov.	Valid	Carlos Rando Harald Pieper Josep A. Alcover Storrs L. Olson	Quaternary		Шаблон:Flag:	A Strigidae.
?Parvelorius calidris	Gen. nov. et Sp. nov.	Valid	Vanessa L. De Pietri Gerald Mayr	Early Miocene	MN 2a		A Scolopacidae, the new genus is erected for the species "Tringa" gracilis Milne-Edwards, 1868, the type species of the new genus and makes a Com. nov. with it; genus may also contain this new species.
Phylloscopus miocaenicus	Sp. nov.	Valid	Jenö Kessler János Hír	Middle Miocene	MN 6-8		A Phylloscopidae.
Picavus litencicensis	Gen. nov. et Sp. nov.	Valid	Gerald Mayr Růžena Gregorová	Early Oligocene	Menilite Formation		A stem group representative of the Piciformes, placed in a new family Picavidae Mayr et Gregorová, 2012. it is the type species of the new genus.
Pipilo naufragus	Sp. nov.	Valid	Storrs L. Olson David B. Wingate	Middle Pleistocene to Holocene	Cave deposits		A Passerellidae, towhee.
Pliocalcarius orkhonensis	Gen. nov. et Sp. nov	Valid	Nikita V. Zelenkov Evgeny N. Kurochkin	Late Pliocene	Villafranchian, Chikoi Formation, MN 16a		Initially considered to be a likely close relative of longspurs, Calcariidae, and made the type species of the new genus Pliocalcarius; Palastrova & Zelenkov (2020) reinterpreted this species as a lark and transferred it to the genus Eremophila.
Praealauda hevesensis	Gen. nov. et Sp. nov.	Valid	Jenö Kessler János Hír	Middle Miocene	MN 7/8		An Alaudidae. The type species of the new genus.
Primophaps schoddei	Gen. nov. et Sp. nov.	Valid	Trevor H. Worthy	Late Oligocene to early Miocene	Riversleigh		A Columbidae related to the bronzewing pigeons. The type species of the new genus.
Pterocnemia mesopotamica	Sp. nov.	Valid	Federico L. Agnolín Jorge I. Noriega	Late Miocene	Ituzaingó Formation, Huayquerian		A Rheidae.
Rallicrex litkensis	Sp. nov.	Valid	Jenö Kessler János Hír	Early Miocene	MN 5		A Rallidae, a new species of Rallicrex Lambrecht, 1933, the type species is Rallicrex kolozsvarensis Lambrecht, 1933.
Rhodospiza shaamarica	Sp. nov.	Valid	Nikita V. Zelenkov Evgeny N. Kurochkin	Late Pliocene	Villafranchian, Chikoi Formation, MN 16a	Шаблон:Flag Шаблон:Flag	Originally described as a species of Rhodospiza; Palastrova & Zelenkov (2020) transferred it to the genus Emberiza.
Sanshuiornis zhangi	Gen. nov. et Sp. nov.	Valid	Wang Min Gerald Mayr Zhang Jiangyong Zhou Zhonghe	Middle Eocene	Huayong Formation		A Ciconiiformes Incertae Sedis.
Schizooura lii	Gen. nov. et Sp. nov	Valid	Zhou Shuang Zhou Zhonghe Jingmai K. O'Connor	Aptian	Jiufotang Formation		A basal Ornithuromorpha. The type species of the new genus.
Scolopacimilis lartetianus	Gen. nov. et Comb. nov.	Valid	Vanessa L. De Pietri Mayr	Early Miocene	MN 2a		A Scolopacidae, a new genus for the species "Totanus" lartetianus, the type species of the genus, creating a Comb. nov.
Shengjingornis yangi	Gen. nov et Sp. nov.	Valid	Li Li Wang Jinqi Zhang Xi Hou Shilin	Early Cretaceous	Jiufotang Formation		A Longipterygidae Zhang, Zhou, Hou L. H. et Gu, 2000, Enantiornithes Walker, 1981. The type species of the new genus.
Songzia acutunguis	Sp. nov.	Valid	Wang Min Gerald Mayr Zhang Jiangyong Zhou Zhonghe	Early Eocene	Yangxi Formation		A Songziidae Hou, 1990, Gruiformes. The type species is Songzia heidangkouensis Hou, 1990.
Sturnus kretzoii	Sp. nov.	Valid	Jenö Kessler János Hír	Late Miocene	MN 9		A Sturnidae.
Tadorna minor	Sp. nov.	Valid	Jenö Kessler János Hír	Miocene	MN 7/8		An Anatidae.
Turdicus minor	Sp. nov.	Valid	Jenö Kessler János Hír	Early to Middle Miocene			A Turdidae.
Tydea septentrionalis	Gen. nov. et Sp. nov.	Valid	Gerald Mayr Thierry Smith	Early Oligocene	Rupelian	Шаблон:Flag:	A Diomedeidae. The type species of the new genus.
Xiangornis shenmi	Gen. nov. et Sp. nov.	Valid	Hu Dongyu Xu Xing Hou Lianhai Corwin Sullivan	Early Cretaceous	Jiufotang Formation		An Enantiornithes Walker, 1981. The type species of the new genus.

Pterosaurs

Research

• A study on the bone histology of Rhamphorhynchus and its implications for inferring life history of members of the genus is published by Prondvai et al. (2012).

New taxa

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Aerotitan	Gen. et sp. nov	Valid	Novas et al.	Late Cretaceous			An azhdarchid. The type species is Aerotitan sudamericanus.
Bellubrunnus	Gen. et sp. nov	Valid	Hone et al.	Late Kimmeridgian	Pfraundorf-Heitzenhofen Basin		A rhamphorhynchid. The type species is Bellubrunnus rothgaengeri.	Файл:Bellubrunnus.png Bellubrunnus.
Dendrorhynchoides mutoudengensis	Sp. nov	Valid	Lü & Hone	Middle Jurassic	Tiaojishan Formation		An anurognathid. Originally described as a species of Dendrorhynchoides, but subsequently transferred to the separate genus Luopterus.
Europejara	Gen. et sp. nov	Valid	Vullo et al.	Upper Barremian	La Huérguina Formation		A tapejarid. The type species is Europejara olcadesorum.	Файл:Europejara olcadesorum.png Europejara.
Gladocephaloideus	Gen. et sp. nov	Valid	Lü, Ji, Wei, & Liu	Barremian	Yixian Formation		A ctenochasmatid ctenochasmatoid.	Файл:Gladocephaloideus jingangshanensis.PNG Life restoration
Guidraco	Gen. et sp. nov	Valid	Wang, Kellner, Jiang, & Cheng	Early Cretaceous	Jiufotang Formation		A close relative of Ludodactylus. The type species is Guidraco venator.	Файл:Guidraco.jpg Guidraco.
Jianchangnathus	Gen. et sp. nov	Valid	Cheng, Wang, Jiang, & Kellner	Middle Jurassic	Tiaojishan Formation		A scaphognathid.
Moganopterus	Gen. et sp. nov	Valid	Lü et al.	Early Cretaceous	Yixian Formation		A boreopterid. The type species is Moganopterus zhuiana.
Qinglongopterus	Gen. et sp.	Valid	Lü, Unwin, Zhao, Gao & Shen.	Middle Jurassic	Tiaojishan Formation		A rhamphorhynchid.

Archosauria incertae sedis

Newly named archosaurs with uncertain affinities

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Samrukia	Gen. et sp. nov	Valid	Naish, Dyke, Cau, Escuillié, Godefroit	Late Cretaceous	Kyzylorda District		A possible basal ornithuromorph; alternatively, it could be a large pterosaur.
Smok	Gen. et sp. nov	Valid	Niedźwiedzki, Sulej & Dzik	Latest Norian/Early Rhaetian	Lipie Śląskie clay-pit		A large predatory archosaur of uncertain phylogenetic placement, the largest known predatory archosaur from the Late Triassic and Early Jurassic of central Europe.