Английская Википедия:2019 in brachiopod paleontology

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Research

• A study aiming to reconstruct the life cycle of the Early Cambrian brachiopods is published by Madison & Kuzmina (2019).
• A study on the anatomy of plectambonitoid specimens from the Middle Ordovician of Russia, and on its implications for the knowledge of life cycles of these brachiopods, is published online by Madison & Kuzmina (2019).
• A study on the petrographic and geochemical preservation of Ordovician dalmanelloid shells from the Lexington Formation of Kentucky, Sheguindah Shale of Ontario and the Stony Mountain Formation of Manitoba, aiming to test the hypothesis of paleo-latitudinal zonation of the shelly benthos, is published by Azmy & Jin (2019).
• A study on the phylogenetic relationships among strophomenoid brachiopods and on the biogeographical changes in the strophomenoids through time (focusing on the impact of the Late Ordovician mass extinction on the evolutionary history of strophomenoids) is published by Congreve, Krug & Patzkowsky (2019).
• A study on the internal structure of the shell of Semiplanella carinthica is published by Pakhnevich (2019), who names a new tribe Semiplanellini in the subfamily Gigantoproductinae.
• A study on the diet of species of Gigantoproductus from the Viséan of Derbyshire (United Kingdom), and on its implications for the knowledge of mechanisms that enabled those brachiopod species to reach large sizes, is published by Angiolini et al. (2019).
• A study on the relative importance of brachiopods and bivalves in the fossil assemblages from the Carboniferous Pennsylvanian Breathitt Formation of Kentucky is published by Hsieh, Bush & Bennington (2019).
• Description of a specimen of a Permian brachiopod species Spiriferella protodraschei bearing ventrally directed spiralia (lophophore‐supporting, coiled internal structures), and a study on the morphology of these structures and their implications for the knowledge of the feeding system of this brachiopod, is published by Lee et al. (2019).
• A study on the evolution of the body size of brachiopods from the Late Permian to the Middle Triassic, as indicated by brachiopod specimens from South China, is published by Chen et al. (2019).
• A study on the biogeography of Pliensbachian brachiopods in western Tethys Ocean is published online by Vörös & Escarguel (2019).
• A study on changes in the body size of benthic marine brachiopods and bivalves from the Lusitanian Basin (Portugal) before the Toarcian oceanic anoxic event is published by Piazza et al. (2019).
• A study on the impact of the early Toarcian extinction event on fossil brachiopods and bivalves known from the Iberian Range (Spain) is published by Danise et al. (2019).

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Aegiria apta	Sp. nov	Valid	Huang et al.	Silurian (Rhuddanian)	Zhangwan Formation
Alaskorhynchus	Gen. et sp. nov	Valid	Baranov & Blodgett	Devonian (Pragian)	Soda Creek Limestone		A member of Rhynchonellida belonging to the family Eatoniidae. The type species is A. sodacreekensis.
Angustothyris dagysi	Sp. nov	Valid	Guo, Chen & Harper	Middle Triassic (Anisian)	Qingyan Formation
Angustothyris qingyanensis	Sp. nov	Valid	Guo, Chen & Harper	Middle Triassic (Anisian)	Qingyan Formation
Anidanthus parvimucronata	Nom. nov	Valid	He & Shen in He et al.	Permian (Changhsingian)	Talung Formation		A member of Productida belonging to the family Linoproductidae and the subfamily Anidanthinae; a replacement name for Anidanthus mucronata He & Shen in He et al. (2005).
Anidanthus subquadratus	Sp. nov	Valid	He, Shi & Shen in He et al.	Permian (Changhsingian)			A member of Productida belonging to the family Linoproductidae and the subfamily Anidanthinae.
Aporthophyla prisca	Sp. nov	Valid	Cocks & Popov	Early Ordovician
Arcullina? enokiani	Sp. nov	Valid	Lee & Shi in Lee et al.	Permian (Kungurian)	Kozhim Formation Kozhim Rudnik Formation		A member of Spiriferida belonging to the family Spiriferellidae.
Askerina	Gen. et sp. nov	Valid	Baarli	Ordovician (Hirnantian) and Silurian (Aeronian)	Solvik		A member of the family Atrypidae. The type species is A. cymbula. Announced in 2019; the final version of the article naming was published in 2021.
Aulacothyris stevensi	Sp. nov	Valid	MacFarlan	Early Jurassic (Aratauran)	Diamond Peak Group		A member of Terebratulida belonging to the family Zeilleriidae.
Aulonotreta neptuni	Sp. nov	Valid	Holmer et al.	Ordovician (Darriwilian )			A member of Lingulida belonging to the family Aulonotretidae.
Australocoelia boucoti	Sp. nov	Valid	De Rezende, Machado & Ponciano	Devonian
Austriellula iordanae	Sp. nov	Valid	Gaetani in Grădinaru & Gaetani	Triassic			A member of Rhynchonellida belonging to the family Norellidae and the subfamily Norellinae.
Bellistrophia	Gen. et comb. nov	Valid	Holmer et al.	Cambrian	Atei Formation Meagher Formation	Шаблон:Flag Шаблон:Flag	A member of Kutorginida. The type species is "Nisusia" deissei Bell (1941); genus also includes "Nisusia" montanensis Bell (1941).
Bernousia	Gen. et sp. nov	Valid	Gourvennec	Paleozoic	Tindouf Basin		A member of Spiriferida. Genus includes new species B. brevedorsata.
Biernatia rhapsody	Sp. nov	Valid	Lavié, Serra & Feltes	Middle and Late Ordovician	Las Aguaditas Formation Las Chacritas Formation		A member of Acrotretida belonging to the family Biernatidae.
Borellithyris	Gen. et sp. nov	Valid	Dulai	Miocene (Tortonian)	Sant’Agata Fossili Formation		A member of Terebratulida belonging to the family Megathyrididae. The type species is B. gaetanii.
Cathaysiorthis xichuanensis	Sp. nov	Valid	Huang et al.	Silurian (Rhuddanian)	Zhangwan Formation
Caucasorhynchella	Gen. nov	Valid	Guo, Chen & Harper	Middle Triassic (Anisian)
Chonosteges cooperi	Sp. nov	Valid	Torres-Martínez et al.	Permian	Paso Hondo Formation
Cleiothyridina pfaffenbergensis	Sp. nov	Valid	Mottequin et al.	Carboniferous (Tournaisian)	Gleitsch Formation		A member of Athyridida belonging to the family Athyrididae.
"Clorinda" symmetrica	Sp. nov	Valid	Cocks	Silurian (Llandovery)
Conotreta andina	Sp. nov	Valid	Lavié, Serra & Feltes	Ordovician (Darriwilian)	Las Aguaditas Formation Las Chacritas Formation		A member of Acrotretida belonging to the family Acrotretidae.
Coolinia diabolica	Sp. nov	Valid	Cocks	Silurian (Llandovery)
Cordatia	Gen. et sp. nov	Valid	Brock & Claybourn in Betts et al.	Early Cambrian			A member of Paterinata possibly belonging to the family Paterinidae. The type species is C. erinae.
Craniops popovi	Sp. nov	Valid	Cocks	Silurian (Llandovery)
Crassispirifer galinae	Sp. nov	Valid	Manankov	Permian			A member of Spiriferida.
Crassumbo germanicus	Sp. nov	Valid	Mottequin et al.	Carboniferous (Tournaisian)	Gleitsch Formation		A member of Spiriferida belonging to the family Crassumbidae.
Cyrtia wrekinensis	Sp. nov	Valid	Cocks	Silurian (Llandovery)
Cyrtina evanescens	Sp. nov	Valid	Gourvennec	Paleozoic	Tindouf Basin		A member of Spiriferinida.
Cyrtospirifer aouinetensis	Sp. nov	Valid	Gourvennec	Paleozoic	Tindouf Basin
Cyrtospirifer (Elasmospirifer)	Subgen. et sp. nov	Valid	Gourvennec	Paleozoic	Tindouf Basin		The subgenus includes new species C. (E.) djebiletensis.
Cyrtospirifer robardeti	Sp. nov	Valid	Gourvennec	Paleozoic	Tindouf Basin
Dictyothyropsis vogli	Sp. nov	Valid	Bujtor & Vörös	Early Cretaceous
Dihelictera askeriensis	Sp. nov	Valid	Baarli	Ordovician (Hirnantian) and Silurian (Aeronian)	Solvik		A member of the family Atrypidae. Announced in 2019; the final version of the article naming was published in 2021.
Dinobolus athelstani	Sp. nov	Valid	Cocks	Silurian (Llandovery)
Eleutherokomma djezairensis	Sp. nov	Valid	Gourvennec	Paleozoic	Tindouf Basin		A member of Spiriferida.
Eospirifer majesticus	Sp. nov	Valid	Cocks	Silurian (Llandovery)
Erbotreta elankii	Sp. nov	Valid	Ushatinskaya & Korovnikov	Cambrian			A member of Acrotretida.
Eucalathis giulioi	Sp. nov	Valid	Dulai	Miocene (Tortonian)	Sant’Agata Fossili Formation		A member of Terebratulida belonging to the family Chlidonophoridae.
Farewellirhynchia	Gen. et sp. nov	Valid	Baranov & Blodgett	Devonian (Pragian)	Soda Creek Limestone		A member of Rhynchonellida belonging to the subfamily Leiorhynchinae. The type species is F. kulkovi.
Fascistropheodonta? wiltzensis	Sp. nov	Valid	Jansen	Early Devonian	Wiltz Formation	Шаблон:Flag Шаблон:Flag	A member of the family Strophodontidae.
Felsithyris	Gen. et sp. nov	Valid	Mottequin & Weyer	Carboniferous (Mississippian)			A member of Spiriferida. The type species is F. hercynica.
Garciaalcaldia	Gen. et sp. nov	Valid	Baranov & Blodgett	Devonian (Pragian)	Soda Creek Limestone		A member of Rhynchonellida belonging to the subfamily Glossinunilinae. The type species is G. alaskensis.
Ghorispirifer	Gen. et comb. nov	Valid	Mottequin & Brice	Devonian (Famennian)		Шаблон:Flag Шаблон:Flag Шаблон:Flag	A Cyrtiopsinae. The type species is "Cyrtiopsis graciosa" chakhaensis Brice (1971) (raised to the rank of a separate species G. chakhaensis), genus also includes "Cyrtiopsis" lapparenti Brice (1971).
Hedeinopsis africana	Sp. nov	Valid	Gourvennec	Paleozoic	Tindouf Basin		A member of Spiriferida.
Howellella minsterleyensis	Sp. nov	Valid	Cocks	Silurian (Llandovery)
Ingentistrophia	Gen. et comb. nov	Valid	Jansen	Early Devonian	Katzenelnbogen Formation Seifen Formation	Шаблон:Flag Шаблон:Flag	A member of Orthotetida belonging to the family Areostrophiidae. The type species is "Orthothethes" ingens Drevermann (1904).
Jonesea oepiki	Sp. nov	Valid	Cocks	Silurian (Llandovery)
Jordanithyris	Gen. et sp. nov	Valid	Feldman et al.	Middle Jurassic (Bathonian and Callovian)	Hamam Mughanniyya		A member of Terebratulida. Genus includes new species J. ardainensis. Announced in 2019; the final version of the article naming it was published in 2021.
Joviatrypa nakremi	Sp. nov	Valid	Baarli	Silurian (Aeronian)	Solvik		A member of the family Atrypidae. Announced in 2019; the final version of the article naming was published in 2021.
Judinica	Gen. et 2 sp. et comb. nov	Valid	Oleneva	Devonian		Шаблон:Flag Шаблон:Flag	A member of Spiriferida belonging to the family Adolfiidae. The type species is J. siratchoica; genus also includes new species J. pseudodeflexa, as well as J. pseudomultifida (Vandercammen, 1955), J. biverrucosa (Vandercammen, 1955) and J. solita (Ljaschenko, 1959).
Koeveskallina bifurcata	Sp. nov	Valid	Guo, Chen & Harper	Middle Triassic (Anisian)	Qingyan Formation
Kosoidea australis	Sp. nov	Valid	Zabini et al.	Ordovician (Hirnantian)	Iapó Formation Vila Maria Formation		A member of Discinoidea.
Kutchithyris simoni	Sp. nov	Valid	Feldman et al.	Middle Jurassic (Callovian)	Mughanniyya		Announced in 2019; the final version of the article naming it was published in 2021.
Lingulipora budili	Sp. nov	Valid	Mergl	Middle Devonian	Srbsko Formation		A member of the family Lingulidae.
Linnarssonia ulakhani	Sp. nov	Valid	Ushatinskaya & Korovnikov	Cambrian			A member of Acrotretida.
Linoporella maxima	Sp. nov	Valid	Cocks	Silurian (Llandovery)
Loboidothyris fordycei	Sp. nov	Valid	MacFarlan	Early Jurassic (Lower Aratauran to Upper Ururoan)		Шаблон:Flag Шаблон:Flag	A member of Terebratulida belonging to the family Loboidothyrididae.
Lobothyris simesi	Sp. nov	Valid	MacFarlan	Early Jurassic (Aratauran to Ururoan)			A member of Terebratulida belonging to the family Lobothyrididae.
Martinia georgei	Sp. nov	Valid	Tazawa in Tazawa, Osawa & Nagura	Carboniferous (Viséan–Bashkirian)	Karaumedate Formation	Шаблон:Flag Шаблон:Flag Шаблон:Flag Шаблон:Flag Шаблон:Flag Шаблон:Flag	A member of Spiriferida belonging to the family Martiniidae.
Martinia liuqiaoiensis	Sp. nov	Valid	He, Shi & Shen in He et al.	Permian (Changhsingian)	Talung Formation		A member of Spiriferida belonging to the family Martiniidae.
Menarhynchus	Gen. et sp. nov	Valid	Baranov & Blodgett	Devonian (Pragian)	Soda Creek Limestone		A member of Rhynchonellida belonging to the subfamily Sphaerirhynchinae. The type species is M. kuskokwimensis.
Micromartinia	Gen. et sp. nov	Junior homonym	He & Weldon in He et al.	Permian (Changhsingian)	Talung Formation		A member of Spiriferida belonging to the superfamily Martinioidea. The type species is M. kwangsiella. The generic name is preoccupied by Micromartinia Amsel (1957).
Minutomarginifera	Nom. nov	Valid	He et al.	Carboniferous and Permian		Шаблон:Flag Шаблон:Flag	A member of Strophalosiidina belonging to the family Chonopectidae; a replacement name for Eileenella Racheboeuf in Wongwanich et al. (2004).
Minutomena missa	Sp. nov	Valid	Huang et al.	Ordovician (Hirnantian)	Kuanyinchiao Beds
Multispirifer kropotkini	Sp. nov	Valid	Gourvennec	Paleozoic	Tindouf Basin		A member of Spiriferida.
Neocyrtina xui	Sp. nov	Valid	Guo, Chen & Harper	Middle Triassic (Anisian)	Qingyan Formation
Neoliothyrina nakremi	Sp. nov	Valid	Bitner in Hryniewicz et al.	Late Paleocene	Basilika Formation		A member of Terebratulida belonging to the family Sellithyridae.
Neoplatystrophia paludis	Sp. nov	Valid	Cocks	Silurian (Llandovery)
Neospirifer borealis	Sp. nov	Valid	Manankov	Permian			A member of Spiriferida.
Nisusia multicostata	Sp. nov	Valid	Holmer et al.	Cambrian (Drumian)	Mila Formation		A member of Kutorginida.
Nottina	Gen. et sp. nov	Valid	Baarli	Silurian (Rhuddanian and Aeronian)	Solvik		A member of the family Atrypidae. The type species is N. phalerata. Announced in 2019; the final version of the article naming was published in 2021.
Nucleospira elgari	Sp. nov	Valid	Cocks	Silurian (Llandovery)
Nudirostralina minuta	Sp. nov	Valid	Guo, Chen & Harper	Middle Triassic (Anisian)	Qingyan Formation
Opsiconidion nanus	Sp. nov	Valid	Mergl	Devonian (Eifelian)	Acanthopyge Limestone		A member of Acrotretoidea belonging to the family Biernatidae.
Oretanomena	Gen. et sp. nov.	Valid	Reyes-Abril, Villas & Gutierrez-Marco	Ordovician (Darriwilian)			A member of Strophomenoidea. Genus includes new species O. meloui.
Ortarhynchia	Gen. et sp. nov	Valid	Gaetani in Grădinaru & Gaetani	Triassic			A member of Rhynchonellida belonging to the family Norellidae and the subfamily Paranorellininae. The type species is O. petersi.
Pachyschizophoria amygdalina	Sp. nov	Valid	Jansen	Early Devonian	Berlé Formation Emsquarzit Formation Hohenrhein Formation	Шаблон:Flag Шаблон:Flag	A member of Enteletoidea belonging to the family Schizophoriidae.
Parabrekia	Gen. et sp. nov	Valid	Guo, Chen & Harper	Middle Triassic (Anisian)	Qingyan Formation		Genus includes new species P. yangi.
Paurorthis? llangynogensis	Sp. nov	Valid	Cocks & Popov	Early Ordovician
Pentameroides johnsoni	Sp. nov	Valid	Cocks	Silurian (Llandovery)
Permorhipidomella	Gen. et sp. nov	Valid	He, Shi & Shen in He et al.	Permian (Changhsingian)			A member of Orthida belonging to the family Rhipidomellidae. Genus includes new species P. ovatus.
Piarorhynchella kittli	Sp. nov	Valid	Gaetani in Grădinaru & Gaetani	Triassic			A member of Rhynchonellida belonging to the family Norellidae and the subfamily Holcorhynchellinae.
Platytrochalos (Salopicoelia) morrellswoodensis	Subgen. et sp. nov	Valid	Cocks	Silurian (Llandovery)
Productina saalfeldensis	Sp. nov	Valid	Mottequin et al.	Carboniferous (Tournaisian)	Gleitsch Formation Hangenberg Limestone		A member of Productida belonging to the family Productellidae.
Ptychomentzelia dobrogeana	Sp. nov	Valid	Gaetani in Grădinaru & Gaetani	Triassic			A member of Spiriferida belonging to the family Mentzeliidae.
Ptychomentzelia simionescui	Sp. nov	Valid	Gaetani in Grădinaru & Gaetani	Triassic			A member of Spiriferida belonging to the family Mentzeliidae.
Quiringites bitami	Sp. nov	Valid	Gourvennec	Paleozoic	Tindouf Basin		A member of Spiriferida.
Rhipidomella parvula	Sp. nov	Valid	He, Shi & Shen in He et al.	Permian (Changhsingian)			A member of Orthida belonging to the family Rhipidomellidae.
Roemerithyris	Gen. et comb. nov	Valid	Mottequin & Weyer	Carboniferous (Mississippian)			A member of Spiriferida. The type species is "Spirifer" macrogaster Roemer (1852).
Rugithyris hasibuani	Sp. nov	Valid	MacFarlan	Early Jurassic (Upper Ururoan)			A member of Terebratulida belonging to the superfamily Loboidothyridoidea.
Rutorhynchia? trigonalis	Sp. nov	Valid	Guo, Chen & Harper	Middle Triassic (Anisian)	Qingyan Formation
Schachriomonia spiraensis	Sp. nov	Valid	Baarli	Ordovician-Silurian	Solvik		A member of the family Atrypidae. Announced in 2019; the final version of the article naming was published in 2021.
Schellwienella justinianoi	Sp. nov	Valid	Rezende, Ponciano & Brett	Late Devonian	Longá Formation
Sellithyris elizabetha	Sp. nov	Valid	Rojas & Sandy	Early Cretaceous (Valanginian)	Rosablanca Formation
Septatrypa salopiensis	Sp. nov	Valid	Cocks	Silurian (Llandovery)
Shelvothyris	Gen. et comb. nov	Valid	Cocks	Silurian (Llandovery)			Genus includes S. furcata (Sowerby, 1839)
Sifella	Gen. et sp. nov	Valid	Baarli	Silurian (Aeronian)	Solvik		A member of the family Atrypidae. The type species is S. patera. Announced in 2019; the final version of the article naming was published in 2021.
Skenidoides pontyfennensis	Sp. nov	Valid	Cocks & Popov	Early Ordovician
Smirnovina ferraria	Sp. nov	Valid	Bujtor & Vörös	Early Cretaceous
Spiriferella protodraschei	Sp. nov	Valid	Lee & Shi in Lee et al.	Permian (late Artinskian–early Kungurian)	Kapp Starostin Formation		A member of Spiriferida belonging to the family Spiriferellidae.
Tamarirhynchia	Gen. et comb. nov	Valid	Baranov & Blodgett	Devonian (Pragian)	Soda Creek Limestone		A member of Rhynchonellida belonging to the subfamily Hebetoechiinae. The type species is "Lancemyonia" varia Tcherkesova (1969).
Tegulithyris? plencnerae	Sp. nov	Valid	MacFarlan	Early Jurassic (Upper Aratauran and Ururoan)			A member of Terebratulida belonging to the family Tegulithyrididae.
Thebesia shelvensis	Sp. nov	Valid	Cocks	Silurian (Llandovery)
Thuringorhynchus	Gen. et sp. nov	Valid	Mottequin et al.	Carboniferous (Tournaisian)	Gleitsch Formation Hangenberg Limestone		A member of Rhynchonellida belonging to the family Pugnacidae. The type species is T. pseudoequitans.
Tornquistia changhsingia	Sp. nov	Valid	He, Shi & Shen in He et al.	Permian (Changhsingian)			A member of Chonetidina belonging to the family Anopliidae and the subfamily Anopliinae.
Trigonithyris wilsoni	Sp. nov	Valid	Feldman et al.	Middle Jurassic (Callovian)	Mughanniyya		Announced in 2019; the final version of the article naming it was published in 2021.
Triplesia habberleyensis	Sp. nov	Valid	Cocks	Silurian (Llandovery)
Tritoechia bolohaulensis	Sp. nov	Valid	Cocks & Popov	Early Ordovician
Tumarinia uldsiensis	Sp. nov	Valid	Manankov	Permian			A member of Spiriferinida.
Ukhtomica	Gen. et sp. nov	Valid	Oleneva	Devonian			A member of Spiriferida belonging to the family Echinospiriferidae. The type species is U. lata; genus also includes U. apschakensis (Krylova, 1955).
Virgianoides	Gen. et comb. nov	Valid	Jin, Mikulic & Kluessendorf	Silurian (Rhuddanian)	Mayville Dolomite	Шаблон:Flag (Шаблон:Flag)	A member of Pentamerida belonging to the family Virgianidae. The type species is BVirgiana barrandei var. major Savage (1916), raised to the rank of a separate species V. major.
Weiningia ziyunensis	Sp. nov	Valid	Yuan et al.	Carboniferous (Serpukhovian)	Baizuo Formation		A member of Spiriferida belonging to the family Martiniidae.
Whitfieldella alvarezensis	Sp. nov	Valid	Cocks	Silurian (Llandovery)
Xenocrania	Gen. et comb. nov	Valid	Chen & Rong	Ordovician		Шаблон:Flag Шаблон:Flag Шаблон:Flag Шаблон:Flag Шаблон:Flag?	The type species is "Palaeocyclus" haimei Reed.
Zeilleria recessa	Sp. nov	Valid	MacFarlan	Early Jurassic (Hettangian)			A member of Terebratulida belonging to the family Zeilleriidae.
Zeilleria sacciformis	Sp. nov	Valid	MacFarlan	Early Jurassic (Ururoan)	Ururoa Formation		A member of Terebratulida belonging to the family Zeilleriidae.
Zeilleria terezowae	Sp. nov	Valid	MacFarlan	Early Jurassic (Aratauran and Ururoan)	Ururoa Formation		A member of Terebratulida belonging to the family Zeilleriidae.
Zittelina hofmanni	Sp. nov	Valid	Bujtor & Vörös	Early Cretaceous
Zygospiraella nupera	Sp. nov	Valid	Baarli	Silurian (Aeronian)	Solvik		A member of the family Atrypidae. Announced in 2019; the final version of the article naming was published in 2021.
Zygospiraella? ventricosa	Sp. nov	Valid	Cocks	Silurian (Llandovery)