Английская Википедия:2022 in paleobotany
Шаблон:ToC Right Шаблон:Year nav topic20 This paleobotany list records new fossil plant taxa that were to be described during the year 2022, as well as notes other significant paleobotany discoveries and events which occurred during 2022.
Algae
Charophytes
Name | Novelty | Status | Authors | Age | Unit | Location | Synonymized taxa | Notes | Images |
---|---|---|---|---|---|---|---|---|---|
Sp. nov |
In press |
Feist & Floquet |
Late Cretaceous |
A charophyte. |
|||||
Sp. nov |
In press |
Feist & Floquet |
Late Cretaceous |
A charophyte. |
|||||
Sp. nov |
In press |
Feist & Floquet |
Late Cretaceous |
A charophyte. |
Charophyte research
- A study on the Paleocene charophyte flora from the South Gobi area in the Junggar Basin (China) and on the Paleogene fossil record of charophytes is published by Cao et al. (2022), who interpret their findings as evidence of the dispersal of charophyte lineages from Asia to Europe in the middle to late Eocene, possibly facilitated by waterbirds.[2]
Chlorophytes
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et comb. nov |
Valid |
Grgasović |
Middle Triassic |
A green alga belonging to the group Dasycladales. Genus includes "Oligoporella" karrerioidea Pia (1935). |
||||
Gen. et sp. nov |
LoDuca in LoDuca et al. |
A green alga belonging to the group Bryopsidales. Genus includes new species E. fredricksi. |
||||||
Sp. nov |
In press |
Torromé & Schlagintweit |
A green alga belonging to the group Dasycladales. |
|||||
Gen. et comb. nov |
Valid |
Grgasović |
Late Triassic, possibly also Middle Triassic |
A green alga belonging to the group Dasycladales. Genus includes "Diplopora" lotharingica Benecke (1898), "Physoporella" jomdaensis Flügel & Mu (1982) and "Physoporella" zamparelliae Parente & Climaco (1999). |
||||
Gen. et sp. nov |
Chai, Aria & Hua |
A green alga belonging to the family Codiaceae. Genus includes new species P. sinense. |
||||||
Sp. nov |
Valid |
Vachard & Krainer |
Permian-Triassic transition |
A green alga belonging to the group Dasycladales. |
Lycopodiopsida
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Valid |
Deng in Deng et al. |
Tongchuan Formation |
A member of the family Isoetaceae. |
||||
Gen. et sp. nov |
Valid |
Herrera et al. |
Huolinhe Formation |
A member of the family Lycopodiaceae. Genus includes new species L. oellgaardii. |
||||
Gen. et sp. nov |
Valid |
Edwards, Li & Berry |
Early Devonian |
A member of Protolepidodendrales of uncertain phylogenetic placement. Genus includes new species M. sinense. |
||||
Sp. nov |
Spiekermann, Jasper, Guerra-Sommer & Uhl in Spiekermann et al. |
A member of Lycopodiopsida of uncertain affinities. |
||||||
Gen. et sp. nov |
Liu et al. |
Wutong Formation |
A member of Isoetales of uncertain affinities. Genus includes new species O. gigas. |
|||||
Sp. nov |
Deng in Deng et al. |
Middle Triassic (Ladinian) |
A lycopsid. |
|||||
Gen. et sp. nov |
Valid |
Prestianni et al. |
An isoetalean lycopsid. Genus includes new species P. minitensis. |
|||||
Sp. nov |
Li & Wang in Li et al. |
Cretaceous (Albian-Cenomanian) |
A species of Selaginella. |
|||||
Sp. nov |
In press |
Li et al. |
Cretaceous |
Burmese amber |
A species of Selaginella. |
Lycopsid research
- Description of new fossil material of Guangdedendron micrum, providing new information on the morphology of this plant, is published by Gao et al. (2022).[17]
- Xu, Liu & Wang (2022) describe new fossil material of Sublepidodendron grabaui from the Devonian (Famennian) Wutong Formation (China), providing new information on the morphology of the female reproductive organs of this plant.[18]
Marchantiophyta
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Valid |
Feldberg, Schäfer-Verwimp & Renner in Feldberg et al. |
Cretaceous (Albian-Cenomanian) |
Burmese amber |
A liverwort, a species of Radula. |
|||
Sp. nov |
Valid |
Feldberg, Schäfer-Verwimp & Renner in Feldberg et al. |
Cretaceous (Albian-Cenomanian) |
Burmese amber |
A liverwort, a species of Radula. |
|||
Sp. nov |
Valid |
Santos et al. |
A liverwort belonging to the family Ricciaceae. |
|||||
Ricciopsis cortaderitaensis[21] |
Gen. et sp. nov |
Valid |
Savoretti et al. |
Middle Triassic |
A liverwort. |
|||
Sp. nov |
Valid |
Savoretti et al. |
Middle Triassic |
A liverwort. |
Marchantiophyta research
- New specimens of Radula heinrichsii, providing new information on the morphology of this liverwort, are described from the Cretaceous Burmese amber by Wang et al. (2022).[22]
Ferns and fern allies
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Valid |
Skog & Sender |
A member of the family Hymenophyllaceae. |
|||||
Sp. nov |
In press |
Trevisan et al. |
Late Cretaceous |
|||||
Sp. nov |
In press |
Zhou et al. |
Early Permian |
A botryopteid fern. |
||||
Gen. et 2 sp. nov |
In press |
Pšenička et al. |
A leptosporangiate fern. Genus includes new species D. chlupatum and D. zlatkokvacekii. |
|||||
Sp. nov |
In press |
Ren & Sun in Ren et al. |
A fern |
|||||
Sp. nov |
Cantrill et al. |
Miocene |
A species of Gleichenia. |
|||||
Sp. nov |
Li & Wang in Li et al. |
Cretaceous (Albian-Cenomanian) |
Burmese amber |
A member of the family Hymenophyllaceae. Originally described as a species of Hymenophyllites, but subsequently moved to the genus Trichomanes sensu lato by Li et al. (2023).[29] |
||||
Sp. nov |
Li & Wang in Li et al. |
Cretaceous (Albian-Cenomanian) |
Burmese amber |
A member of the family Hymenophyllaceae. |
||||
Sp. nov |
Li & Wang in Li et al. |
Cretaceous (Albian-Cenomanian) |
Burmese amber |
A member of the family Hymenophyllaceae. |
||||
Sp. nov |
Long, Wang & Shi in Long et al. |
Cretaceous |
A species of Microlepia. |
|||||
Gen. et sp. nov |
Valid |
Nishida et al. |
Late Cretaceous |
A probable stem polypod leptosporangiate fern. Genus includes new species M. rothwellii. |
||||
Gen. et sp. nov |
Valid |
Morales-Toledo, Mendoza-Ruiz & Cevallos-Ferriz |
A member of the family Dicksoniaceae. Genus includes new species P. jurassica. |
|||||
Sp. nov |
Barbosa et al. |
Douro Carboniferous Basin |
A member of Equisetales. |
|||||
Sp. nov |
In press |
Zhang et al. |
Early Permian |
Taiyuan Formation |
A member of Marattiales belonging to the family Psaroniaceae. |
|||
Nom. nov |
Valid |
Deshmukh |
A member of Stauropteridales; a replacement name for Gillespiea Erwin & Rothwell (1989). |
Fern and fern ally research
- Pecopteris lativenosa is interpreted as a member of the late Paleozoic marattialean family Psaroniaceae by Li et al. (2022).[36]
Gnetales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
Valid |
Manchester et al. |
Late Jurassic |
Morrison Formation |
A gnetale. Genus includes the species B. hoodiorum. |
|||
Gen. et sp. nov |
In press |
Ren et al. |
Early Cretaceous |
Chijinbao Formation |
A member of the family Ephedraceae. Genus includes new species D. beishanensis. |
Bennettitales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Valid |
Saadatnejad |
Late Triassic (Rhaetian) |
A member of Bennettitales. |
||||
Sp. nov |
Valid |
Saadatnejad |
Late Triassic (Rhaetian) |
Kalariz Formation |
A member of Bennettitales. |
|||
Gen. et sp. nov |
Valid |
Pott & Takimoto |
A member of Bennettitales. Genus includes new species K. densifolia. |
|||||
Sp. nov |
Valid |
Kvaček |
Late Cretaceous (Cenomanian) |
Ginkgophytes
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Sun & Deng in Sun et al. |
Late Triassic |
Шаблон:Flag | |||||
Glossophyllum panii[42] |
Sp. nov |
Sun & Deng in Sun et al. |
Late Triassic |
Шаблон:Flag | ||||
Sp. nov |
In press |
Nosova & Kostina |
||||||
Sp. nov |
In press |
Dong et al. |
Middle-Late Jurassic |
|||||
Sp. nov |
In press |
Dong et al. |
Middle-Late Jurassic |
Daohugou Beds |
Ginkgophyte research
- Revision of Ginkgo abaniensis, based on data from leaves from the Jurassic Mura Formation (Russia), is published by Frolov & Mashchuk (2022), who emend the diagnosis of this species, and transfer Ginkgo abaniensis, Ginkgo glinkiensis and Ginkgo capillata to the genus Ginkgoites.[45]
Conifers
Araucariaceae
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Valid |
Bodnar et al. |
Cheirolepidiaceae
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Cheng et al. |
A probable member of the family Cheirolepidiaceae. |
||||||
Sp. nov |
In press |
Mendes & Kvaček |
Figueira da Foz Formation |
A member of the family Cheirolepidiaceae. |
||||
Sp. nov |
Kvaček & Mendes |
Early Cretaceous (Aptian-Albian) |
Figueira da Foz Formation |
Cupressaceae
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Valid |
Sadowski, Schmidt & Kunzmann |
Eocene |
Europe (Baltic Sea region) |
Cupressaceous pollen cone. |
|||
Gen. et sp. nov |
Valid |
Andruchow-Colombo et al. |
A member of the family Cupressaceae. Genus includes new species P. lefipanensis. |
Pinaceae
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Shi, Sun, Meng & Yu in Shi et al. |
Early Cretaceous (Albian) |
A Keteleeria-like wood morphogenus. |
|||||
Sp. nov |
Li & Dong in Dong et al. |
Miocene |
Changtai Formation |
A species of Nothotsuga. |
||||
Sp. nov |
Bazhenova, Wu & Jin in Bazhenova et al. |
Late Pleistocene |
Maoming Basin |
A pine. |
||||
Sp. nov |
Li, Hu & Xiao in Li et al. |
Miocene |
Shengxian Formation |
A pine. |
Podocarpaceae
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Announced |
Pujana et al. |
Oligocene |
San José Formation |
A podocarpaceous wood morphospecies |
|||
Sp. nov |
Announced |
Pujana et al. |
Oligocene |
San José Formation |
A podocarpaceous wood morphospecies |
|||
Sp. nov |
Castañeda-Posadas |
Miocene |
A species of Podocarpus. |
Sciadopityaceae
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
In press |
Jiang et al. |
Middle to Late Jurassic (Callovian to Kimmeridgian) |
A member of the family Sciadopityaceae. Genus includes new species Z. yanliaoense. |
Voltziales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
Announced |
Wang et al. |
Permian (Cisuralian) |
Shanxi Formation |
A voltzialean conifer. |
Other conifers
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
Barbacka & Górecki in Barbacka et al. |
A needle leaf similar to the leaves of Pinus. Genus includes new species A. triangulatum. |
||||||
Sp. nov |
In press |
Cai, Zhang & Feng in Cai et al. |
Late Permian |
|||||
Gen. et sp. nov |
In press |
Forte, Kustatscher & Van Konijnenburg-van Cittert in Forte et al. |
Genus includes new species S. tridentata. |
|||||
Gen. et comb. nov |
Valid |
Kerp et al. |
Permian |
The type species is "Lebachia" laxifolia (1939); |
Conifer research
- Bodnar et al. (2022) reassess the anatomy and systematics of the permineralized conifer-like woods from the Triassic strata from Argentina, confirm the assignment of the logs related to the families Cupressaceae and Cheirolepidiaceae, as well as three taxa related to Araucariaceae (Agathoxylon cozzoi, Agathoxylon protoaraucana and Agathoxylon argentinum), and argue that the fossil woods previously assigned to the families Podocarpaceae and Taxaceae do not have enough preserved characters to support such assignment.[64]
- A study on the pattern of conifer turnover across the Cretaceous-Paleogene boundary in the Raton and Denver basins (Colorado, United States) is published by Berry (2022).[65]
- Mantzouka, Akkemik & Güngör (2022) describe fossil woods of Cupressinoxylon matromnense from the middle Miocene Eşelek volcanic deposits (Gökçeada, Turkey), preserved with feeding damage produced by members of the agromyzid genus Protophytobia, and supporting the existence of an eastern Mediterranean Miocene Climatic Optimum hotspot which additionally included Greek islands of Lemnos and Lesbos.[66]
Flowering plants
Chloranthales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Valid |
Friis et al. |
Early Cretaceous (Aptian-Albian) |
Almargem Formation |
||||
Gen. et sp. nov |
Valid |
Friis et al. |
Early Cretaceous (Aptian-Albian) |
Almargem Formation |
Genus includes new species P. portugallicus. |
Magnoliids
Laurales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
In press |
Rubalcava-Knoth & Cevallos-Ferriz |
Cretaceous (Albian–Cenomanian) |
La Cintura Formation |
||||
Sp. nov |
Valid |
Vasquez-Loranca & Cevallos-Ferriz |
Miocene |
A member of the family Lauraceae. |
||||
Gen. et sp. nov |
Valid |
Maccracken et al. |
A member of the family Lauraceae. Genus includes new species C. gettyi. |
|||||
Sp. nov |
Valid |
Akkemik, Iamandei & Çelik |
Early Miocene |
Hançili Formation |
Fossil wood of a member of the family Lauraceae. |
|||
Sp. nov |
Valid |
Vasquez-Loranca & Cevallos-Ferriz |
Miocene |
A member of the family Lauraceae. |
||||
Sp. nov |
Valid |
Vasquez-Loranca & Cevallos-Ferriz |
Miocene |
A member of the family Lauraceae. |
||||
Mezilaurinoxylon draconis[69] |
Sp. nov |
Valid |
Vasquez-Loranca & Cevallos-Ferriz |
Miocene |
A member of the family Lauraceae. |
|||
Mezilaurinoxylon miocenica[69] |
Sp. nov |
Valid |
Vasquez-Loranca & Cevallos-Ferriz |
Miocene |
A member of the family Lauraceae. |
Magnoliales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Valid |
Martinetto in Niccolini et al. |
Miocene (Messinian) |
Piedmont Basin |
A species of Magnolia. |
Piperales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
Valid |
Friis, Crane & Pedersen |
Early Cretaceous (Aptian–Albian) |
A member of the family Aristolochiaceae. Genus includes new species A. huberi. |
||||
Gen. et sp. nov |
Valid |
Friis, Crane & Pedersen |
Late Cretaceous (Cenomanian) |
A member of the family Aristolochiaceae. Genus includes new species S. mauldinense. |
Monocots
Lilioid monocots
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Valid |
Rozefelds et al. |
Oligocene |
A species of Pandanus. |
Commelinid monocots
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
In press |
Kumar, Hazra & Khan in Kumar et al. |
Late Cretaceous-Paleocene (Maastrichtian-Danian) |
A member of the family Arecaceae belonging to the subfamily Coryphoideae. |
||||
Sp. nov |
In press |
Kumar, Hazra & Khan in Kumar et al. |
Late Cretaceous-Paleocene (Maastrichtian-Danian) |
Deccan Intertrappean Beds |
A member of the family Arecaceae belonging to the subfamily Coryphoideae. |
Monocot research
- Leaf fossils of costapalmate-palms belonging to the genus Sabalites are described from the ?Santonian–Campanian Belly River Group, Campanian Foremost Formation (Alberta, Canada) and Maastrichtian Frenchman Formation (Saskatchewan, Canada) by Greenwood, Conran & West (2022), who interpret the studied fossils as constraining climate reconstructions for the Late Cretaceous high mid-latitudes of North America (c. 55° N) to exclude significant freezing episodes; the authors also transfer the Late Cretaceous species "Geonomites" imperialis to the genus Phoenicites, and reassess Sabalites carolinensis as more likely to be Campanian than Coniacian–Santonian in age.[77]
- A study on the impact of the absence of megaherbivores in the aftermath of the Cretaceous–Paleogene extinction event on the evolution of palms is published by Onstein, Kissling & Linder (2022).[78]
- A study on the evolutionary history of palms belonging to the group Mauritiinae, as inferred from a phylogenetic analysis incorporating fossil data, is published by Bacon et al. (2022).[79]
Basal eudicots
Proteales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
Huegele & Wang |
Early Cretaceous (Albian) |
A platanaceous inflorescence. Genus includes new species D. hoisingtonensis. |
|||||
Gen et sp nov |
Valid |
Huegele & Manchester |
A platanaceous flowering head. |
|||||
Sp. nov |
Moiseeva, Kodrul & Jin |
Late Eocene |
A species of Meliosma. |
|||||
Sp. nov |
Luo & Jia in Luo et al. |
Miocene |
Upper Youshashan Formation |
A species of Nelumbo. |
||||
Sp. nov |
In press |
Dong et al. |
Miocene |
Fotan Group |
A species of Nelumbo. |
|||
Comb nov |
in press |
(Wolfe) |
Puget Group |
A platanaceous leaf. |
||||
Comb nov |
in press |
(Chen) |
Eocene |
A platanaceous leaf. |
||||
Sp. nov |
Golovneva et al. |
Early Cretaceous (Albian) |
A member of the family Platanaceae. |
|||||
Sapindopsis orientalis[86] |
Sp. nov |
Golovneva et al. |
Early Cretaceous (Albian) |
A member of the family Platanaceae. |
Protealean research
- Redescription of the Okanagan Highlands genus Langeria with description of associated stipules and reproductive structures plus formal reassignment of the genus to Platanaceae by Huegele & Manchester is published.[81]
Ranunculales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Valid |
Denk & Sami in Denk et al. |
Pleistocene (Calabrian) |
A species of Berberis. |
||||
Sp. nov |
Tang et al. |
Pliocene |
A species of Mahonia. |
|||||
Sp. nov |
Wu & Zhou in Wu et al. |
Eocene |
Shuanghe Formation |
A member of the family Menispermaceae. |
Superasterids
Aquifoliales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Nom. nov |
Valid |
Doweld |
Late Cretaceous (Maastrichtian) |
A holly; a replacement name for Ilex antiqua Knobloch & Mai (1986). |
||||
Nom. nov |
Valid |
Doweld |
Miocene (Messinian) |
A holly; a replacement name for Ilex myricoides Massalongo (1858). |
Caryophyllales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
In press |
Estrada-Ruiz |
Miocene |
A species of Podopterus. |
Cornales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Nom. nov |
Valid |
Deshmukh |
Pleistocene |
A member of the family Cornaceae; a replacement name for Cornoxylon Blackwell (1982). |
||||
Gen. et comb. nov |
Valid |
Manchester & Collinson |
Early Eocene |
A mastixioid fruit; a new genus for Beckettia mastixioides Reid & Chandler (1933). |
Dipsacales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
In press |
Huang & Zhou in Huang et al. |
Late Pliocene |
Heqing Basin |
A species of Sambucus. |
Ericales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
In press |
Koutecký & Sakala in Koutecký, Sakala & Chytrý |
Ústí Formation |
A member of the family Ebenaceae. Genus includes new species P. kvacekii. |
Icacinales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Atkinson |
Late Cretaceous (Campanian) |
A member of the family Icacinaceae. |
Metteniusales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Estrada-Ruiz et al. |
Miocene |
A species of Calatola. |
Superrosids
Cucurbitales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
In press |
Huegele & Manchester |
Paleocene |
A member of the family Cucurbitaceae. Genus includes new species L. potamoglossensis. |
Fabales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Announced |
Nguyen, Su & J. Huang in Nguyen et al. |
Miocene |
Yen Bai Basin |
An Albizia species. |
|||
Sp. nov |
Valid |
Ramos et al. |
Late Pleistocene |
El Palmar Formation |
A member of the family Fabaceae. |
|||
Sp. nov |
Valid |
Ramos et al. |
Late Pleistocene |
El Palmar Formation |
A species of Cedrelinga. |
|||
Sp. nov |
Valid |
Akkemik, Iamandei & Çelik |
Early Miocene |
Hançili Formation |
Fossil wood of a member of the family Fabaceae. |
|||
Gen. et sp. nov |
Valid |
Ramos et al. |
Late Pleistocene |
El Palmar Formation |
A member of the family Fabaceae. Genus includes new species C. yukeriense. |
|||
Sp. nov |
Valid |
Ramos et al. |
Late Pleistocene |
El Palmar Formation |
A member of the family Fabaceae. |
|||
Sp. nov |
Announced |
Bhatia, Srivastava & Mehrotra |
Late Paleocene |
Tura Formation |
A fabaceous seed pod morphospecies. |
|||
Sp. nov |
Valid |
Ramos et al. |
Late Pleistocene |
El Palmar Formation |
A member of the family Fabaceae. |
|||
Gen. et comb. nov |
Valid |
Herendeen et al. |
Latest Paleocene to late early Eocene |
A member of Papilionoideae; a new genus for "Acacia" lamarensis Knowlton (1899). |
||||
Gen. et sp. nov |
Valid |
Ramos et al. |
Late Pleistocene |
El Palmar Formation |
A Fabaceae genus. |
|||
Sp. nov |
Announced |
Bhatia, Srivastava & Mehrotra |
Late Paleocene |
Tura Formation |
A fabaceous leaf morphospecies. |
|||
Sp. nov |
In press |
Li, Huang & Su in Li et al. |
Late Eocene |
Lunpola Basin |
A member of the family Fabaceae. |
|||
Gen. et sp. nov |
Valid |
Ramos et al. |
Late Pleistocene |
El Palmar Formation |
A member of the family Fabaceae. Genus includes new species P. uniseriatum. |
|||
Gen. et comb. nov |
Valid |
Herendeen et al. |
Eocene |
A member of Papilionoideae; a new genus for "Diplotropis" claibornensis Herendeen & Dilcher (1990). |
Fabalean research
- New fossil material of members of the genus Bauhinia is described from the Eocene of the Puyang Basin (China) by Jia et al. (2022), who interpret their findings as the earliest reliable fossil records of Bauhinia in Asia.[104]
- Moya et al. (2022) study the affinities of fossil legumes Entrerrioxylon victoriensis, Gossweilerodendroxylon palmariensis, Paraoxystigma concordiensis and Cylicodiscuxylon paragabunensis from the Cenozoic Paraná, Arroyo Feliciano and El Palmar formations (Argentina) with extant West African legumes, and discuss the possible migration routes by which these plants may have arrived in South America from Africa.[105]
Fagales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
In press |
Wang et al. |
Miocene |
Fotan Group |
A species of Castanopsis. |
|||
Sp. nov |
Xiao & Ji in Ji et al. |
Miocene |
Hannuoba Formation |
A species of Comptonia. |
||||
Sp. nov |
Valid |
Akkemik, Iamandei & Çelik |
Early Miocene |
Fossil wood of a member of the family Myricaceae. |
||||
Sp. nov |
Announced |
Pujana et al. |
Oligocene |
San José Formation |
A nothofagaceous wood morphospecies |
|||
Sp. nov |
Valid |
Stults, Tiffney & Axsmith |
Pliocene |
A species of Pterocarya. |
||||
Sp. nov |
In press |
Liu & Jin in Liu et al. |
Late Oligocene |
Yongning Formation |
An oak. |
|||
Sp. nov |
In press |
Liu & Jin in Liu et al. |
Late Oligocene |
Yongning Formation |
An oak. |
|||
Sp. nov |
In press |
Liu & Jin in Liu et al. |
Late Oligocene |
Yongning Formation |
An oak. |
|||
Sp. nov |
In press |
Liu & Jin in Liu et al. |
Late Oligocene |
Yongning Formation |
An oak. |
Malpighiales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et comb. nov |
Valid |
Hamersma et al. |
Early Oligocene |
A member of the family Euphorbiaceae; a new genus for "Jatropha" tertiara Berry. |
||||
Sp. nov |
Valid |
Bennike in Bennike et al. |
Probably early Pleistocene |
A species of Elatine. Announced in 2022; the final article version was published in 2023. |
||||
Sp. nov |
In press |
Akkemik et al. |
Late Oligocene-Early Miocene |
İstanbul Formation |
A Mammea relative wood morphospecies |
|||
Comb. nov |
In press |
(Nelson & Jud) Akkemik & D. Mantzouka |
Miocene |
A Mammea relative wood morphospecies |
||||
Sp. nov |
Valid |
Grote in Grote, Duangkrayom & Jintasakul |
Late Miocene |
Tha Chang beds |
A species of Parinari. |
|||
Sp. nov |
Valid |
Grote in Grote, Duangkrayom & Jintasakul |
Late Miocene |
Tha Chang beds |
A species of Parinari. |
|||
Sp. nov |
In press |
Miocene |
A species of Plukenetia. |
Malvales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Manchester et al. |
Late Cretaceous-Paleocene (Maastrichtian–Danian) |
A member of the family Malvaceae. |
|||||
Malvacipolloides intertrappea[115] |
Sp. nov |
Manchester et al. |
Late Cretaceous-Paleocene (Maastrichtian–Danian) |
Deccan Intertrappean Beds |
A member of the family Malvaceae. |
|||
Sp. nov |
Valid |
Hazra, Mahato & Khan in Hazra et al. |
A species of Thespesia. |
Malvalean research
- A study on the evolutionary history of Dipterocarpaceae, as indicated by biogeography of pollen fossils from Africa and India, molecular data and fossil amber records, is published by Bansal et al. (2022).[117]
Myrtales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Dong et al. |
Miocene |
A member of the family Lythraceae belonging to the subfamily Trapoideae. |
|||||
Sp. nov |
Announced |
Pujana et al. |
Oligocene |
San José Formation |
A myrtaceous wood morphospecies. |
|||
Sp. nov |
Han & Jia in Han et al. |
Late Eocene |
Bailuyuan Formation |
|||||
Sp. nov |
Cai et al. |
Miocene |
A water caltrop. |
Oxalidales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp nov |
In press |
Tang, Smith, & Atkinson |
A Cunoniaceous species. |
|||||
Gen. et sp. nov |
Valid |
Baas et al. |
Cretaceous |
A probable Connaraceous wood morphotaxon. |
||||
Gen. et sp. nov |
Valid |
Matel et al. |
Early Eocene |
A member of Cunoniaceae. |
||||
Gen. et sp. nov |
Valid |
Matel et al. |
Early Eocene |
Huitrera Formation |
A member of Cunoniaceae. |
|||
Sp. nov |
Announced |
Pujana et al. |
Oligocene |
San José Formation |
A cunoniaceous wood morphospecies. |
Oxalidalean research
Tand, Smith, and Atkinson describe the first North American instance of the previously Paleo-Antarctic Rainforest Lineage Cunoniaceae fruits from Sucia Island. Previously considered solely a Gondwanan family, the new species indicate a complex geographic history for the group.[121]
Rosales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
Valid |
Shi et al. |
Cretaceous |
A rhamnaceous floral morphotaxon. |
||||
Sp. nov |
Valid |
El-Noamani |
A member of the family Moraceae. |
|||||
Sp. nov |
In press |
Dong et al. |
Miocene |
A species of Ficus. |
||||
Sp. nov |
In press |
Dong et al. |
Miocene |
A species of Ficus. |
||||
Sp. nov |
Valid |
Shi et al. |
Cretaceous |
Burmese amber |
A species of Phylica. |
|||
Sp. nov |
Hazra et al. |
Pliocene |
A species of Ventilago. |
|||||
Sp. nov |
Hazra et al. |
Miocene |
A species of Ventilago. |
Sapindales
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et comb. nov |
Valid |
Manchester & Judd |
Eocene |
An anacardiaceous samara. |
||||
Sp. nov |
In press |
Yin et al. |
Miocene |
A species of Canarium. |
||||
Sp. nov |
Xiang & Jin in Xiang et al. |
Late Pleistocene |
Maoming Basin |
A species of Canarium. |
||||
Sp. nov |
Xiao & Wu in Xiao et al. |
Miocene |
Shengxian Formation |
A species of Choerospondias. |
||||
Sp. nov |
Xiao & Wu in Xiao et al. |
Miocene |
Shengxian Formation |
A species of Choerospondias. |
||||
Gen. et sp. nov |
Huang, Morley & Hoorn in Huang et al. |
Eocene |
A member of the family Sapindaceae. Genus includes new species G. burmanica. |
|||||
Sp. nov |
Valid |
Chen, Del Rio & Su in Chen et al. |
Eocene |
Niubao Formation |
A species of Koelreuteria. |
|||
Gen. et sp. nov |
In press |
Manchester & Judd |
Eocene |
Ione Formation |
A member of the family Anacardiaceae. Genus includes new species L. weeksae. |
|||
Gen. et comb. nov |
Valid |
Strullu-Derrien et al. |
Eocene |
A fruit of likely sapindalean affinity; a new genus for "Juglandicarya" gruetii Vaudois-Miéja (1976). |
Other Eudicots
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
In press |
Rothwell & Stockey |
Late Cretaceous |
St. Mary River Formation |
An aquatic eudicot. |
Other angiosperms
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Gen. et sp. nov |
Chen & Wang |
A flower bud of an early angiosperm. Genus includes new species A. lingyuanensis. |
||||||
Gen. et sp. nov |
Announced 2022 |
Čepičková & Kvaček |
A Basal angiosperm leaf morphogenus |
|||||
Gen. et sp. nov |
Valid |
Heřmanová et al. |
An angiosperm fruit of uncertain affinities, with similarities to the family Dilleniaceae. |
|||||
Gen. et sp. nov |
Valid |
Friis et al. |
Early Cretaceous (Aptian-Albian) |
Almargem Formation |
A flowering plant of uncertain position at the level of ANA-grade angiosperms-Chloranthaceae-magnoliids. Genus includes new species E. paisii. |
|||
Gen. et sp. nov |
Valid |
Friis et al. |
Early Cretaceous (Aptian-Albian) |
Almargem Formation |
A flowering plant of uncertain position at the level of ANA-grade angiosperms-Chloranthaceae-magnoliids. Genus includes new species E. cateficensis. |
|||
Sp. nov |
Du et al. |
Early Cretaceous |
A herbaceous eudicot. |
|||||
Gen. et sp. nov |
In press |
Cui et al. |
Middle-Late Jurassic |
A possible flower bud. |
||||
Gen. et sp. nov |
Valid |
Wu et al. |
Early Cretaceous (Aptian) |
Chijinpu Formation |
An early flowering plant. Genus includes new species G. beishanensis. |
|||
Gen. et sp. nov |
In press |
Frolov & Enushchenko |
Irkutsk Coal Basin |
An angiosperm with leaf epidermal structure most similar to those of modern Asparagales and Liliales. Genus includes new species H. antiqua. |
||||
Gen. et sp. nov |
In press |
Huegele & Manchester |
Paleocene |
Fort Union Formation |
A flowering plant of uncertain affinities. Genus includes new species H. bighornensis. |
|||
Gen. et sp. nov |
Valid |
Friis et al. |
Early Cretaceous (Aptian-Albian) |
Almargem Formation |
A flowering plant of uncertain position at the level of ANA-grade angiosperms-Chloranthaceae-magnoliids. Genus includes new species I. cuneiformis. |
|||
Gen. et sp. nov |
Wang |
Early Cretaceous (Barremian–Aptian) |
Yixian Formation |
An early angiosperm. Genus includes new species L. inexpectus. |
||||
Gen. et 2 sp. nov |
Gobo et al. |
Originally described as a member or a relative of the family Ranunculaceae, but subsequently considered to be a mesangiosperm of uncertain affinities, possibly a magnoliid.[147] Genus includes new species S. lobata and S. acuta. |
||||||
Comb. nov |
Announced |
(Velenovský) Čepičková & Kvaček |
A Basal angiosperm leaf morphogenus |
|||||
Gen. et sp. nov |
Valid |
Edmonds, Stockey & Rothwell |
Late Cretaceous (Maastrichtian) |
An aquatic dicot. Genus includes new species T. aquatica. |
||||
Gen. et sp. nov |
Valid |
Friis et al. |
Early Cretaceous (Aptian-Albian) |
Almargem Formation |
A flowering plant of uncertain position at the level of ANA-grade angiosperms-Chloranthaceae-magnoliids. Genus includes new species V. globiferus. |
General angiosperm research
- Surangea mohgaoensis, originally interpreted as fern megaspores, is reinterpreted as angiosperm fruits by Ramteke et al. (2022).[149]
- Zhang et al. (2022) describe rich assemblages of spiny plant fossils from the Eocene (Bartonian) Niubao Formation (Tibet, China), preserving seven different spine morphologies, and interpret this finding as evidence of the presence of a diversity of spiny plants in Eocene central Tibet, as well as evidence of a rapid diversification of spiny plants in Eurasia around that time.[150]
- A preliminary report on a new fossil angiosperm flora of the Lesvos Petrified Forest at Akrocheiras east of Sigri on Lesbos, Greece is given by Kafetzidou et al. Preliminary taxa identifications are given and commentary on the climactic implications are made.[151]
- A study aiming to determine the relationship between past atmospheric CO2 and temperature fluctuations and the shifts in diversification rates of Poaceae and Asteraceae is published by Palazzesi et al. (2022).[152]
Other plants
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
In press |
Nosova & Tekleva |
Prisayan Formation |
Pollen cone with pollen of ginkgoalean or gnetophytalean affinity. |
||||
Gen et sp nov |
Announced |
Pujana et al. |
Oligocene |
San José Formation |
A wood morphospecies of uncertain affinity. |
|||
Sp. nov |
Valid |
Frolov, Kazanovsky & Enushchenko |
Early Jurassic (Toarcian) |
Middle Subformation of Prisayan Formation |
A member of Bryopsida of uncertain affinities. |
|||
Gen. et sp. nov |
Santos & Wang |
A cone-like reproductive organ of a seed plant. Genus includes new species C. triassica. |
||||||
Sp. nov |
Valid |
Gomankov |
Permian |
A member of Cycadales. |
||||
Sp. nov |
In press |
Conceição et al. |
Pedra de Fogo Formation |
A gymnosperm. |
||||
Gen. et sp. nov |
Shi et al. |
Early Cretaceous |
A seed-bearing structure of a corystosperm seed fern. Genus includes new species J. zhoui. |
|||||
Sp. nov |
Barbacka in Barbacka et al. |
Early Jurassic (Hettangian) |
Zagaje Formation |
Cuticle of a seed fern. |
||||
Sp. nov |
In press |
Correia et al. |
Douro Carboniferous Basin |
An early gymnosperm. |
||||
Sp. nov |
Valid |
Frolov, Kazanovsky & Enushchenko |
Early Jurassic (Pliensbachian) |
Lower Subformation of Prisayan Formation |
A member of Bryopsida of uncertain affinities. |
|||
Gen. et sp. nov |
In press |
Ma et al. |
Permian (Wuchiapingian) |
A gigantopterid. Genus includes new species P. qingloongensis. |
||||
Sp. nov |
Wang et al. |
A euphyllophyte of uncertain affinities. |
||||||
Gen. et sp. nov |
Valid |
Naugolnykh |
Ordovician |
A rhyniophyte of uncertain affinities. The type species is P. caudatum. |
||||
Gen. et sp. nov |
In press |
Conceição et al. |
Permian (Cisuralian) |
Pedra de Fogo Formation |
A gymnosperm. Genus includes new species P. merlottii. |
|||
Sp. nov |
Colston, Landaw & Tomescu |
Devonian (Emsian) |
A member of the group Trimerophytopsida. |
|||||
Gen. et sp. nov |
In press |
Friis, Crane & Pedersen |
Early Cretaceous (Albian) |
A seed plant similar to Brenneria potomacensis. Genus includes new species R. zhoui. |
||||
Gen. et sp. nov |
In press |
Krings |
Early Devonian |
An alga, probably a green alga belonging to the family Mesotaeniaceae. Genus includes new species R. velatum. |
||||
Gen. et sp. nov |
Valid |
Zhang et al. |
Linjia Formation |
A member of Glossopteridales. The type species is S. sunii. |
||||
Gen. et sp. nov |
Valid |
Naugolnykh & Mogutcheva |
Keshinskian/Keshinskaya Formation |
A member of Peltaspermales belonging to the family Angaropeltaceae. Genus includes new species T. triassica. |
||||
Gen. et sp. nov |
Valid |
McSweeney, Shimeta & Buckeridge |
Norton Gully Sandstone Formation |
A plant of uncertain affinities, similar to members of Zosterophyllopsida. Genus includes new species T. garrattii. |
||||
Gen. et sp. nov |
In press |
Huang, Liu & Xue |
Mangshan Group |
A polysporangiate land plant. Genus includes new species T. antrorsa. |
||||
Gen. et sp. nov |
Valid |
Rothwell et al. |
Jurassic |
A stem-cycad. Genus includes new species T. maahlae. |
||||
Sp. nov |
In press |
Li et al. |
Cretaceous |
Burmese amber |
A moss belonging to the group Hypnodendrales. |
|||
Gen. et sp. nov |
Valid |
Gensel |
Devonian (Emsian) |
A basal euphyllophyte. Genus includes new species W. phocarum. |
||||
Gen. et sp. nov |
Valid |
Klymiuk, Rothwell & Stockey |
Early Cretaceous (Valanginian) |
A gymnosperm of uncertain phylogenetic placement, possibly having affinities with gnetophytes or angiosperms. Genus includes new species X. quatsinoensis. |
||||
Gen. et sp. nov |
Yang & Wang |
Devonian (Famennian) |
A fern-like plant of uncertain affinities. Genus includes new species X. spina. |
Other plant research
- A study on the xylem development in Leptocentroxyla, and on its implications for the knowledge of the evolution of pith, is published by Tomescu & McQueen (2022).[175]
- Decombeix et al. (2022) report evidence of tylosis formation in permineralized wood of Dameria hueberi from the Tournaisian of Australia.[176]
- The first comprehensive crown reconstruction of Medullosa stellata var. typica, based on data from a specimen from the Chemnitz petrified forest (Germany), is presented by Luthardt et al. (2022).[177]
- Fossil material of Rhabdotaenia is reported from the Permian Umm Irna Formation (Jordan) by Blomenkemper et al. (2022), representing the northernmost occurrence of this Gondwanan leaf type reported to date.[178]
Palynology
Name | Novelty | Status | Authors | Age | Type locality | Location | Notes | Images |
---|---|---|---|---|---|---|---|---|
Sp. nov |
Valid |
Playford |
Carboniferous (Mississippian) |
A trilete spore. |
||||
Gen. et sp. nov |
Valid |
Playford |
Carboniferous (Mississippian) |
Lyall Formation |
A trilete spore. Genus includes new species A. cancellosus. |
|||
Sp. nov |
Valid |
Playford |
Carboniferous (Mississippian) |
Lyall Formation |
A trilete spore. |
|||
Camptotriletes suggrandis[179] |
Sp. nov |
Valid |
Playford |
Carboniferous (Mississippian) |
Lyall Formation |
A trilete spore. |
||
Sp. nov |
Valid |
Playford |
Carboniferous (Mississippian) |
Lyall Formation |
A trilete spore. |
|||
Sp. nov |
Valid |
Playford |
Carboniferous (Mississippian) |
Lyall Formation |
A trilete spore. |
|||
Sp. nov |
Valid |
Playford |
Carboniferous (Mississippian) |
Lyall Formation |
A trilete spore. |
|||
Sp. nov |
Valid |
Playford |
Carboniferous (Mississippian) |
Lyall Formation |
A trilete spore. |
|||
Sp. nov |
In press |
Sui, McLoughlin & Feng in Sui et al. |
Permian (Wuchiapingian–Changhsingian) |
Xuanwei Formation |
A lycophyte megaspore. |
|||
Sp. nov |
Hu et al. |
Early Cretaceous |
Bongor Basin |
A gymnosperm pollen. |
||||
Gen. et sp. nov |
In press |
Narváez et al. |
Paleocene (Danian) |
Salamanca Formation |
Pollen of a flowering plant. |
|||
Gen. et sp. nov |
In press |
Legrand, Yamada & Nishida |
Late Cretaceous (Cenomanian–Turonian) |
Mikasa Formation |
A Normapolles-type flowering plant pollen. Genus includes new species Y. mikasaensis. |
Research
- Review of the studies on the origin of the land flora is published by Bowman (2022).[184]
- A study on the evolution of body plans of members of Viridiplantae, based on a review of the fossil record, molecular data and developmental biology, is published by Niklas & Tiffney (2022).[185]
- A study on the biodiversity of land plants at the equator during their first major diversification in the Late Silurian–Early Devonian is published by Wellman et al. (2022).[186]
- A study on the evolution of heterospory during the Devonian is published by Leslie & Bonacorsi (2022).[187]
- Seven coniferous nurse logs that have been colonized by conifer and equisetalean roots are reported from four Permian intervals in the Ordos Basin (China) by Feng et al. (2022), indicating that conifer tree stems probably functioned as hosts to both conspecific and interspecific seedlings in the Cathaysian Flora.[188]
- A study on the impact of the Intertropical Convergence Zone in the emerging South Atlantic region on Aptian plant communities from eight Brazilian sedimentary basins is published by Carvalho et al. (2022), who report evidence of an overall predominance of xerophytic plants, attesting to more dry conditions, and of a humidification trend towards the end of the late Aptian resulting in the predominance of hydrophytes, hygrophytes, tropical lowland flora and upland flora, indicative of prevalence of lowland and montane rainforests.[189]
- A study on the distribution and relative abundances of major plant groups from the Albian Gates Formation (Alberta, Canada) is published by Kalyniuk et al. (2022).[190]
- A study on the relationship between whole-genome duplication, seed traits and the selectivity of the survival of plants during the Cretaceous–Paleogene extinction event is published by Berry & Jaganathan (2022).[191]
- New Oligocene flora is described from the Dong Ho Formation (Vietnam) by Huang et al. (2022), who interpret the studied fossils as evidence of long-term environmental, floristic and vegetational stability in this region since the Paleogene.[192]
- Gentis et al. (2022) describe fossil wood specimens from the Miocene Natma Formation (Myanmar), representing an assemblage dominated by members of the families Fabaceae and Dipterocarpaceae, interpreted as coming from different types of low altitude forest ecosystems (tropical wet evergreen, tropical dry and deciduous, and tropical littoral), and interpreted as indicative of a monsoonal climate with an alternance of a dry season and a wet season.[193]
- Abundant compression floras dominated by angiosperm leaves are described from two sites of probable Pliocene age in Brunei by Wilf et al. (2022), who interpret these floras as evidence of dipterocarp-dominated lowland rainforests in the Malay Archipelago before the Pleistocene.[194]
- A study on the impact of the extinct Neotropical megafauna on the variability in plant functional traits and biome geography in Central and South America is published by Dantas & Pausas (2022).[195]
- A study on plant material from rock overhangs from mid-late Holocene sites along the Kawarau-Cromwell-Roxburgh Gorges in Central Otago (New Zealand), much of which was likely transported as roosting material or consumed by moa birds, and on its implications for the knowledge of the mid-late Holocene regional vegetation of Central Otago and the knowledge of vegetation changes since mid-late Holocene, is published by Pole (2022).[196]
- A study on the role of hydraulic failure in the evolution of early vascular plants is published by Bouda et al. (2022), suggesting that drought selection played a key role in the diversification of vascular arrangements beginning with the Devonian explosion.[197]
References
- ↑ 1,0 1,1 1,2 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 3,0 3,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 15,0 15,1 15,2 15,3 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 19,0 19,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 21,0 21,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 39,0 39,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 42,0 42,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 44,0 44,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 56,0 56,1 56,2 56,3 56,4 56,5 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 60,0 60,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 67,0 67,1 67,2 67,3 67,4 67,5 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 69,0 69,1 69,2 69,3 69,4 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 72,0 72,1 72,2 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 74,0 74,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 76,0 76,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 81,0 81,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 85,0 85,1 Шаблон:Cite journal
- ↑ 86,0 86,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 90,0 90,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 98,0 98,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 100,0 100,1 100,2 100,3 100,4 100,5 100,6 Шаблон:Cite journal
- ↑ 101,0 101,1 Шаблон:Cite journal
- ↑ 102,0 102,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 109,0 109,1 109,2 109,3 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 112,0 112,1 Шаблон:Cite journal
- ↑ 113,0 113,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 115,0 115,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 121,0 121,1 Шаблон:Cite journal
- ↑ 122,0 122,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 124,0 124,1 Шаблон:Cite journal
- ↑ 125,0 125,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 127,0 127,1 Шаблон:Cite journal
- ↑ 128,0 128,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 132,0 132,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 139,0 139,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite book
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 154,0 154,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 157,0 157,1 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ 179,0 179,1 179,2 179,3 179,4 179,5 179,6 179,7 Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal