Английская Википедия:Agave shawii

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Agave shawii is a species of monocarpic succulent plant in the genus Agave, commonly known as Shaw's agave.[1][2] It is a rosette-forming plant characterized by glossy, green leaves with toothed margins. After several years of slow growth, the plant puts all of its resources to produce a towering stalk of flowers, and then dies. The death of the flowering rosette is compensated by the growth of numerous clonal pups. This species is segregated into two subspecies, one native to the coast of southwestern California[3] and northwestern Baja California, known commonly as the coast agave, and another native to the Baja California desert, known as the Goldman agave.[4]

For centuries, this species proved to be an invaluable source of accessible and abundant food for the indigenous peoples in the region, like the Kumeyaay, Tiipai and Paipai. After the European colonization of the Americas, the Spanish missionaries discouraged use of the agave, and moved the native peoples inland.[5] Further development of the agave's habitat by American settlers contributed to the dwindling population of the species. Only two small populations are left within the political boundaries of the United States,[6] one with only a single individual. In Mexico, it is still abundant but threatened by coastal development. Subspecies shawii is designated as an endangered species by the California Native Plant Society and the IUCN.[7][8] The plant is named for Henry Shaw, the founder of the Missouri Botanical Garden.[9]

Description

Agave shawii is a very slow-growing, small-to-medium-sized agave.

Файл:Agave shawii 177375689.jpg
The colorful spines on the margins of the leaves

The foliage is arranged in a rosette that measures Шаблон:Convert to Шаблон:Convert wide. There may be numerous rosettes on top of erect to decumbent trunks that emerge from the rootstock. The foliage is glossy, colored light to dark green, and positioned in an ascending fashion. The leaves are shaped narrow to ovate, measuring Шаблон:Convert long by Шаблон:Convert wide, with the abaxial (lower surface) surface convex. Leaves feel thick, fleshy, and rigid, and their margins are armed with colorful and well-defined spines. The spine at the tip of the leaf measures Шаблон:Convert.[10][11]

The panicle-like inflorescence grows Шаблон:Convert tall. The bracts (leaves on the inflorescence) are persistent, shaped lanceolate to triangular, and are Шаблон:Convert large. Subtended by the bracts are 8–14 lateral umbels (branches), which are Шаблон:Convert long. Each umbel consists of a mass of yellowish or reddish flowers, with 35–75 flowers per cluster. After flowering, it produces fruits Шаблон:Convert large, containing fertilized (black-colored) and unfertilized (white-colored) seeds.[10][11]

It generally flowers September to May, and as typical for agaves, the rosette dies thereafter. Although capable of reproducing by suckering, populations vary considerably in their behavior, with some consisting entirely of individual rosettes, while others form groups or colonies of clones.

Subdivisions

The infraspecific taxa are variously labeled as varieties or subspecies, depending on the source.[3][7][8][11]

  • Agave shawii subsp. shawii — The autonymous subspecies. The southernmost "typical" plants that can easily be distinguished occur near San Quintin. Near El Rosario, both subspecies become difficult to discern.[11]
  • Agave shawii subsp. goldmaniana (Trel.) Gentry — Commonly known as the Goldman agave. The rosette leaves are Шаблон:Convert long, and Шаблон:Convert wide, shaped lanceolate. The inflorescence has 18–25 umbels on a 3–5 meter stem. The long and broader shape of the inflorescence, along with the narrow leaves set it apart from the typical subspecies. It predominates in the coast and desert of the southern part of Baja California state.[4] This subspecies has much higher rates of seed set and seedling recruitment compared to subsp. shawii.[11]

Agave sebastiana, a plant native to Cedros Island and the Vizcaino Peninsula, was formerly placed in this species as Agave shawii var. sebastiana.[12]

Distribution and habitat

This species is primarily distributed in, and near-endemic to, the state of Baja California in Mexico. It was formerly widespread along the coast of San Diego County, California, but coastal development has reduced the entire population to just two natural occurrences, of which only one is wholly natural. There is a Шаблон:Convert north-to-south gap between the southernmost natural plants in San Diego County and the northernmost natural plants in Baja California.[11] The coastal subspecies shawii is the taxon present in San Diego County, and ranges south to El Rosario in Baja California. At El Rosario, it is subsumed by the desert subspecies goldmaniana, which ranges from there to Santa Rosaliíta in southern Baja California.[12]

Subspecies shawii only occurs in a vegetation community known as maritime succulent scrub, which is found along the Pacific Coast of northern Baja California state of Mexico and southwesternmost San Diego County of California. This habitat only extends a few kilometers into the United States, occurring in narrow bands on coastal bluffs with almost constant exposure to coastal winds with high salt content. Maritime succulent scrub is regarded as the most xeric of the coastal sage vegetation types, and is dominated by other succulent plants such as cacti (Bergerocactus, Cylindropuntia, Ferocactus, Mammilaria, Opuntia), cliff spurge (Euphorbia misera) and Crassulacean plants like Dudleya.[13] Subspecies shawii was probably once found north of Torrey Pines, but with the development of all viable habitat, has all but disappeared from the American side of the border.[11]

Subspecies goldmaniana grows in the southern portion of the maritime succulent scrub and in the central desert of southern Baja California.[4]

Uses

Cultivation

Agave shawii is cultivated as an ornamental plant, by specialty plant nurseries. It is used in cactus and succulent gardens, containers,[14] and for drought tolerant and wildlife gardens.[15]

Plants enjoy a sandy loam soil that has good drainage. It is recommended to plant in a location where the sharp spines will not be a hazard. Virtually disease free except for scale insects.[16] Roots are very rapid responders to rain and dry plants start growing feeder (rain) roots after exposure to the rain. Plants develop best color when exposed to full sun along the coast. Some relief from the hot afternoon sun in the inland valleys with partial shade would provide the best results for growers. Within the United States, it is not recommended to grow this plant in desert locations like Las Vegas, but it will thrive in coastal locations from San Diego to San Francisco.[15]

In a preservation effort, Shaw's agaves were introduced into the Torrey Pines State Reserve and Cabrillo National Monument in San Diego, California and have established themselves.[17] There is also large colony of Shaw's Agave in the San Diego Botanic Garden located in Encinitas, California.[18] The San Elijo Lagoon Nature Center in Encinitas also has Shaw's agaves planted as part of their California native plant xeriscaping.[19]

By indigenous peoples

The coastal agave (Tiipai: Шаблон:Lang, Paipai: Шаблон:Lang) was widely utilized by indigenous peoples until the European colonization of the Americas forced natives away from the coastal region where Agave shawii occurs. Subsequently, the lack of interaction between native peoples and their agaves led to the decay of knowledge regarding the usage of the local agaves, with the cultivation of agaves in the entire Kumeyaay-Paipai cultural region now only occurring in the village of Santa Catarina.[20]

The Spanish explorers and missionaries from Sebastián Vizcaíno and Junipero Serra's expeditions noticed the usage of the agave for fiber and food. Serra said on his arrival to the coast near El Descanso, "We saw the land was covered with very good mescal but I think the Indians pay little attention to it due to the abundance of fish and other foods."[21] Serra's observations fail to take into account the seasonal changes in the food supply, as the agave was heavily utilized in other months. Juan Crespí stated that the related Agave deserti was "...a delicious, sweet preserve. This is the wretched heathens’ daily bread."[22]

Agaves had high importance for indigenous food networks, but today the laborious effort of preparing the agaves is rarely undertaken. Preparation of the agave for consumption included locating suitable heads, using a digging stick to extract the heads, constructing an earth oven, where the agave head would be cooked for two days before it was ready to eat. Importance of the agave as a food is supported by archaeological records showing high dental attrition in the remains of indigenous peoples, likely due to the frequent consumption of tough plant fibers from the agave.[5]

The fiber of the agave would have been put to a great variety of uses, for making belts, bowstrings, carrying nets, cordage, sandals, and other indispensable items.[23] Vizcaíno noted that the cordage was used by fishermen to create lines of much higher quality than his own. Some Paipai women still produce agave fiber products today, which are sold as handicrafts, but also offer to teach both natives and non-natives throughout the region.[5][20]

Gallery

See also

Other flora of the maritime succulent scrub:

References

Шаблон:Reflist

  • Raymond M. Turner, Janice E. Bowers, and Tony L. Burgess, Sonoran Desert Plants: an Ecological Atlas (Tucson: The University of Arizona Press, 1995) pp. 63–65

External links

Шаблон:Commons category

Шаблон:Taxonbar

  1. Engelmann, Georg. Transactions of the Academy of Science St. Louis 3:314, t. 2-4. 1875.
  2. Шаблон:Cite journal
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  4. 4,0 4,1 4,2 Шаблон:Cite book
  5. 5,0 5,1 5,2 Wilken, Michael A. (2012) An Ethnobotany of Baja California's Kumeyaay Indians. Retrieved 13 October 2021
  6. Ошибка цитирования Неверный тег <ref>; для сносок NatureServe не указан текст
  7. 7,0 7,1 León de la Luz, J.L. & Vanderplank, S. 2020. Agave shawii subsp. shawii. The IUCN Red List of Threatened Species 2020: e.T125136787A125136804. https://dx.doi.org/10.2305/IUCN.UK.2020-3.RLTS.T125136787A125136804.en. Accessed on 03 February 2022.
  8. 8,0 8,1 Шаблон:Cite web
  9. Starr, Greg. Agaves: living sculptures for landscapes and containers. Timber Press, 2012.
  10. 10,0 10,1 Шаблон:Cite web
  11. 11,0 11,1 11,2 11,3 11,4 11,5 11,6 Шаблон:Cite journal
  12. 12,0 12,1 Шаблон:Cite journal
  13. Шаблон:Cite journal
  14. Шаблон:Cite web
  15. 15,0 15,1 Шаблон:Cite web
  16. Шаблон:Cite web
  17. Ingram, Stephen. Cacti, Agaves, and Yuccas of California and Nevada. Cachuma Press, 2008.
  18. Шаблон:Cite web
  19. Шаблон:Cite web
  20. 20,0 20,1 Wilken-Robertson, M. (2004). Strategies for Sustainable Development of Natural and Cultural Resources in the Paipai Indian Community of Santa Catarina, Baja California. The US-Mexican Border Environment: Tribal Environmental Issues of the Border Region, (9), 71.
  21. Gómez Canedo, Lino (1969) De Mexico a la Alta California: Una Gran Epopoya Misional. Mexico City: Editorial Jus, S.A.
  22. Шаблон:Cite book
  23. Шаблон:Cite web