Английская Википедия:Aizoaceae

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Шаблон:Short description Шаблон:Automatic taxobox

The Aizoaceae (Шаблон:IPAc-en), or fig-marigold family, is a large family of dicotyledonous flowering plants containing 135 genera and about 1800 species.[1] They are commonly known as ice plants or carpet weeds. They are often called vygies in South Africa and New Zealand. Highly succulent species that resemble stones are sometimes called mesembs.

Description

Файл:Mesembryanthemum guerichianum seedling IMG 8167-cropped.jpg
Mesembryanthemum guerichianum seedling, showing the epidermal bladder cells that inspired the name "ice plant".
Файл:LL-Q14196 (afr)-Oesjaar-vygie.wav
Pronunciation of the South African colloquial name, Vygie, for Aizoaceae.

The family Aizoaceae is widely recognised by taxonomists. It once went by the botanical name "Ficoidaceae", now disallowed. The APG II system of 2003 (unchanged from the APG system of 1998) also recognizes the family, and assigns it to the order Caryophyllales in the clade core eudicots. The APG II system also classes the former families Mesembryanthemaceae Fenzl, Sesuviaceae Horan. and Tetragoniaceae Link under the family Aizoaceae.

The common Afrikaans name "vygie" meaning "small fig" refers to the fruiting capsule, which resembles the true fig.[2] Glistening epidermal bladder cells give the family its common name "ice plants".[3]

Most species (96%, 1782 species in 132 genera) in this family are endemic to arid or semiarid parts of Southern Africa in the Succulent Karoo.[4] Much of the Aizoaceae's diversity is found in the Greater Cape Floristic Region, which is the most plant-diverse temperate region in the world.[5] A few species are found in Australia and the Central Pacific area.[6]

Most fig-marigolds are herbaceous, rarely somewhat woody, with sympodial growth and stems either erect or prostrate. Leaves are simple, opposite or alternate, and more or less succulent with entire (or rarely toothed) margins. Flowers are perfect in most species (but unisexual in some), actinomorphic, and appear singularly or in few-flowered cymes developing from the leaf axils. Sepals are typically five (3–8) and more or less connate (fused) below. True petals are absent. However, some species have numerous linear petals derived from staminodes.[7] The seed capsules have one to numerous seeds per cell and are often hygrochastic, dispersing seeds by "jet action" when wet.[3]

Evolution

The radiation of the Aizoaceae, specifically the subfamily Ruschioideae, was one of the most recent among the angiosperms, occurring 1.13–6.49 Mya. It is also one of the fastest radiations ever described in the angiosperms, with a diversification rate of about 4.4 species per million years.[8] This diversification was roughly contemporaneous with major radiations in two other succulent lineages, Cactaceae and Agave.[9]

The family includes many species that use crassulacean acid metabolism as pathway for carbon fixation. Some species in the subfamily Sesuvioideae instead use [[C4 carbon fixation|Шаблон:C4 carbon fixation]], which might have evolved multiple times in the group.[10]

Taxonomy

Файл:Aptenia cordifolia flower leaves.jpg
Aptenia cordifolia or rock rose

Because of the hyperdiversity of the Aizoaceae and the young age of the clade, many generic and species boundaries are uncertain.[8]

Файл:Carpobrotus specimen with pink flowers.jpg
Carpobrotus edulis, an "ice plant"
Файл:Cheiridopsis denticulata.jpg
Cheiridopsis denticulata
Файл:Drosanthemum speciosumP1050020.JPG
Drosanthemum speciosum
Файл:Fenestraria rhopalophylla.jpg
Fenestraria rhopalophylla
Файл:Jensenobotrya lossowiana.jpg
Jensenobotrya lossowiana
Файл:Lithops bromfieldii (25916752581).jpg
Lithops bromfieldii
Файл:Sesuvium p.jpg
Sesuvium portulacastrum

Subfamily Acrosanthoideae

Genera:[11]

Subfamily Aizooideae

Genera:[12]

Subfamily Mesembryanthemoideae

Genera:[13]

Subfamily Ruschioideae

Genera:

Tribe Apatesieae[14]
Tribe Dorotheantheae[15]

* no longer recognised

Tribe Ruschieae[16]

Шаблон:Div col

Шаблон:Div col end

Subfamily Sesuvioideae

This subfamily includes a number of Шаблон:C4 species.[10]

Genera:[18]

Unplaced genera

Include;

Uses

Файл:Starr 050222-4149 Tetragonia tetragonioides.jpg
Tetragonia tetragonoides ("New Zealand spinach")

Several genera are cultivated. Lithops, or "living stones", are popular as novelty house plants because of their stone-like appearance.

Some species are edible, including:

C. edulis was introduced to California in the early 1900s to stabilize soil along railroad tracks and has become invasive.[21] In southern California, ice plants are sometimes used as firewalls;[22] however, they do burn if not carefully maintained.[23]

References

Шаблон:Reflist

Further reading

External links

Шаблон:Commons category

Шаблон:Angiosperm families Шаблон:Taxonbar Шаблон:Authority control