Английская Википедия:Commiphora
Шаблон:Short description Шаблон:Automatic taxobox
The genus of the myrrhs, Commiphora, is the most species-rich genus of flowering plants in the frankincense and myrrh family, Burseraceae. The genus contains approximately 190 species of shrubs and trees, which are distributed throughout the (sub-) tropical regions of Africa, the western Indian Ocean islands, the Arabian Peninsula, India, and South America.[1][2][3] The genus is drought-tolerant and common throughout the xerophytic scrub, seasonally dry tropical forests, and woodlands of these regions.
The common name myrrh refers to several species of the genus, from which aromatic resins are derived for various fragrance and medicinal uses by humans.
Description
Leaves in Commiphora are pinnately compound (or very rarely unifoliolate). Many species are armed with spines. Bark is often exfoliating, peeling in thin sheets to reveal colorful, sometimes photosynthetic, bark below. Stems are frequently succulent, especially in species native to drier environments. Flowers are typically dioecious (subdioecious) and fruits are drupes, usually with a 2-locular ovary (one is abortive).[4] In response to wounding, the stems of many species will exude aromatic resins.
Ecology and biogeography
Commiphora can serve as a model genus for understanding plant evolution in the drier regions of the Old World tropics, particularly in eastern continental Africa and Madagascar, where diversity in the genus is concentrated. The closely related sister genus to Commiphora, Bursera, has been used as a model genus to study patterns of evolution in the New World seasonally dry tropical forests.[5]
Use by humans
Products from many species of Commiphora have been used for various purposes, sometimes as timber, building material, and natural fencing, but more often valued for the aromatic resins produced by several members of the genus. "Myrrh", the common name for these dried resins, is fragrant and has been used both as fragrance and for medicinal purposes (e.g., Balsam of Mecca, C. gileadensis).[6] Use of myrrh resin is frequent and pronounced throughout historical texts of cultural significance, including the Bible.
Systematics and taxonomy
Recent studies using DNA sequence data have confirmed the monophyly of Commiphora;[2][7] however, this data suggests that previous classification of the genus into sections does not reflect monophyletic interspecific relationships.
Species
181 species are accepted:[3]
- Commiphora acuminata Шаблон:Small
- Commiphora africana (A.Rich.) Engl. (syn. Heudelotia africana),[8] sometimes identified with ancient bdellium. Used indirectly by the San bushmen to poison their arrow tips for hunting [9]
- Commiphora alata Шаблон:Small
- Commiphora alaticaulis J.B.Gillett & Vollesen
- Commiphora anacardiifolia Шаблон:Small
- Commiphora andranovoryensis Шаблон:Small
- Commiphora angolensis Engl., also known as "sand commiphora", growing mainly in Angola and Namibia
- Commiphora angustifoliolata Шаблон:Small
- Commiphora ankaranensis Шаблон:Small
- Commiphora antunesii Шаблон:Small
- Commiphora aprevalii Guillaumin, endemic to Madagascar
- Commiphora arafy Шаблон:Small
- Commiphora arenaria Шаблон:Small
- Commiphora baluensis Шаблон:Small
- Commiphora benguelensis Шаблон:Small
- Commiphora berardellii Шаблон:Small
- Commiphora berryi Шаблон:Small
- Commiphora boranensis Vollesen[8]
- Commiphora brevicalyx Шаблон:Small
- Commiphora buruxa Шаблон:Small
- Commiphora caerulea Шаблон:Small
- Commiphora campestris Шаблон:Small
- Commiphora capensis Шаблон:Small
- Commiphora capuronii Шаблон:Small
- Commiphora caudata (Wight & Arn.) Engl.[8]
- Commiphora cervifolia Шаблон:Small
- Commiphora chaetocarpa Шаблон:Small
- Commiphora chevalieri Шаблон:Small
- Commiphora chiovendana Шаблон:Small
- Commiphora ciliata Vollesen
- Commiphora coleopsis Шаблон:Small
- Commiphora confusa Vollesen
- Commiphora corrugata J.B.Gillett & Vollesen[8]
- Commiphora crenatoserrata Шаблон:Small
- Commiphora cuneifolia Шаблон:Small
- Commiphora cyclophylla Шаблон:Small
- Commiphora dalzielii Шаблон:Small
- Commiphora dinteri Шаблон:Small
- Commiphora discolor Шаблон:Small
- Commiphora drake-brockmanii Шаблон:Small
- Commiphora dulcis Шаблон:Small
- Commiphora edulis Шаблон:Small
- Commiphora elliptica Шаблон:Small
- Commiphora eminii Шаблон:Small
- Commiphora engleri Шаблон:Small
- Commiphora enneaphylla Шаблон:Small
- Commiphora erlangeriana Шаблон:Small
- Commiphora erosa Vollesen
- Commiphora falcata Шаблон:Small
- Commiphora foliacea Шаблон:Small
- Commiphora franciscana Шаблон:Small
- Commiphora fraxinifolia Шаблон:Small
- Commiphora fraxinoides Шаблон:Small
- Commiphora fulvotomentosa Шаблон:Small
- Commiphora gardoensis Шаблон:Small
- Commiphora gariepensis Шаблон:Small
- Commiphora giessii Шаблон:Small
- Commiphora gileadensis (L.) C.Chr.[8] (syn. Commiphora opobalsamum), producing balsam of Mecca.
- Commiphora glandulosa Schinz
- Commiphora glaucescens Шаблон:Small
- Commiphora gorinii Шаблон:Small
- Commiphora gracilifrondosa Шаблон:Small
- Commiphora grandifolia Шаблон:Small
- Commiphora grosswelleri Шаблон:Small
- Commiphora guerichiania Шаблон:Small
- Commiphora guidottii Chiov. ex Guid. (syn. Commiphora sessiliflora), producing habak hadi, known as bisabol, opoponax, scented or sweet myrrh.[10]
- Commiphora guillauminii Шаблон:Small
- Commiphora gurreh Шаблон:Small
- Commiphora hartmannii Шаблон:Small
- Commiphora harveyi (Engl.) Engl.
- Commiphora hereroensis Шаблон:Small
- Commiphora hildebrandtii Шаблон:Small
- Commiphora hodai Шаблон:Small
- Commiphora hornbyi Шаблон:Small
- Commiphora horrida Шаблон:Small
- Commiphora humbertii H.Perrier
- Commiphora kaokoensis Шаблон:Small
- Commiphora karibensis Шаблон:Small
- Commiphora kataf (Forssk.) Engl.(syn. Commiphora holtziana Engl, Commiphora erythraea (Ehrenb.) Engl.), producing habak hagar, known as sweet myrrh, sometimes sold as opoponax.[10]
- Commiphora kerstingii Шаблон:Small
- Commiphora kraeuseliana Шаблон:Small
- Commiphora kua (R.Br. ex Royle) Vollesen (syn. Commiphora habessinica (O.Berg) Engl.)
- Commiphora kucharii Шаблон:Small
- Commiphora kuneneana Шаблон:Small
- Commiphora lacerata Шаблон:Small
- Commiphora lamii Шаблон:Small
- Commiphora lasiodisca Шаблон:Small
- Commiphora laxecymigera Шаблон:Small
- Commiphora leandriana Шаблон:Small
- Commiphora leptophloeos (Mart.) J.B.Gillett, the only species of Commiphora present in the Americas.[11]
- Commiphora lobatospathulata Шаблон:Small
- Commiphora longibracteata Шаблон:Small
- Commiphora madagascariensis Jacq.
- Commiphora mafaidoha Шаблон:Small[12]
- Commiphora mahafaliensis Шаблон:Small
- Commiphora marchandii Шаблон:Small
- Commiphora marlothii Шаблон:Small
- Commiphora merkeri Шаблон:Small
- Commiphora merkii Шаблон:Small
- Commiphora mildbraedii Шаблон:Small
- Commiphora mollis Шаблон:Small
- Commiphora mombassensis Шаблон:Small
- Commiphora monoica Vollesen
- Commiphora monstruosa Шаблон:Small
- Commiphora morondavensis Шаблон:Small
- Commiphora mossambicensis (Oliv.) Engl.
- Commiphora mossamedensis Шаблон:Small
- Commiphora mulelame Шаблон:Small
- Commiphora multifoliolata Шаблон:Small
- Commiphora multijuga Шаблон:Small
- Commiphora murraywatsonii Шаблон:Small
- Commiphora myrrha (Nees) Engl. (syn. Commiphora molmol), producing myrrh.
- Commiphora namaensis Шаблон:Small
- Commiphora namibensis Шаблон:Small
- Commiphora neglecta Шаблон:Small
- Commiphora oblanceolata Шаблон:Small
- Commiphora oblongifolia Шаблон:Small
- Commiphora obovata Шаблон:Small
- Commiphora oddurensis Шаблон:Small
- Commiphora omundomba Шаблон:Small
- Commiphora orbicularis Шаблон:Small
- Commiphora ornifolia Шаблон:Small
- Commiphora otjihipana Шаблон:Small
- Commiphora ovalifolia Шаблон:Small
- Commiphora paolii Шаблон:Small
- Commiphora parvifolia Шаблон:Small
- Commiphora pedunculata Шаблон:Small
- Commiphora pervilleana Шаблон:Small
- Commiphora planifrons Шаблон:Small
- Commiphora playfairii Шаблон:Small
- Commiphora pruinosa Шаблон:Small
- Commiphora pseudopaolii Шаблон:Small
- Commiphora pteleifolia Шаблон:Small
- Commiphora pterocarpa Шаблон:Small
- Commiphora pyracanthoides Шаблон:Small
- Commiphora quadricincta Шаблон:Small
- Commiphora quercifoliola Шаблон:Small
- Commiphora rangeana Шаблон:Small
- Commiphora razakamalalae Шаблон:Small
- Commiphora rostrata Шаблон:Small
- Commiphora ruquietiana Шаблон:Small
- Commiphora ruspolii Шаблон:Small
- Commiphora samharensis Шаблон:Small
- Commiphora sarandensis Шаблон:Small
- Commiphora saxicola Engl., Rock corkwood, a shrub endemic to Namibia[13]
- Commiphora schimperi (O.Bergman) Engl.
- Commiphora schlechteri Шаблон:Small
- Commiphora schultzei Шаблон:Small
- Commiphora sennii Шаблон:Small
- Commiphora serrata Шаблон:Small
- Commiphora serrulata Шаблон:Small
- Commiphora setulifera Шаблон:Small
- Commiphora simplicifolia H.Perrier[14]
- Commiphora sinuata Шаблон:Small
- Commiphora socotrana (Balf.f.) Engl.
- Commiphora spathulata Шаблон:Small
- Commiphora spathulifoliolata Шаблон:Small
- Commiphora sphaerocarpa Chiov
- Commiphora spinulosa Шаблон:Small
- Commiphora staphyleifolia Шаблон:Small
- Commiphora stellatopubescens Шаблон:Small
- Commiphora stellulata Шаблон:Small
- Commiphora steynii Шаблон:Small
- Commiphora stocksiana (Engl.) Engl., known in Pakistan as bayisa gugal
- Commiphora sulcata Шаблон:Small
- Commiphora swynnertonii Шаблон:Small
- Commiphora tenuipetiolata Шаблон:Small
- Commiphora tetramera Шаблон:Small
- Commiphora truncata Шаблон:Small
- Commiphora tsimanampetsae Шаблон:Small
- Commiphora ugogensis Шаблон:Small
- Commiphora ulugurensis Шаблон:Small
- Commiphora unilobata J.B.Gillett & Vollesen
- Commiphora viminea Шаблон:Small
- Commiphora virgata Шаблон:Small
- Commiphora wightii (Arn.) Bhandari (syn. Commiphora mukul[15]), producing gum guggul, sometimes identified with ancient bdellium.
- Commiphora wildii Шаблон:Small
- Commiphora woodii Шаблон:Small
- Commiphora zanzibarica Шаблон:Small
Gallery
References
External links
Шаблон:Taxonbar Шаблон:Authority control
- ↑ Daly et al. 2011. Burseraceae. Families and genera of vascular plants. 10:76–104.
- ↑ 2,0 2,1 Weeks, A. and Simpson, B.B. 2007. Molecular phylogenetic analysis of Commiphora (Burseraceae) yields insight on the evolution and historical biogeography of an “impossible” genus. Molecular Phylogenetics and Evolution. 42:62–79.
- ↑ 3,0 3,1 Шаблон:Cite web
- ↑ Gillett, J.B. 1991. Burseraceae. In: Polhill, R.M. (Ed.), Flora of Tropical East Africa. A.A. Balkema, Rotterdam. Pp. 1–95.
- ↑ De Nova, A. et al. 2011. Insights into the historical construction of species-rich Mesoamerican seasonally dry tropical forests: the diversification of Bursera (Burseraceae, Sapindales). New Phytologist. 193(1):276–287.
- ↑ Musselman, L.J. 2007. Figs, Dates, Laurel, and Myrrh: Plants of the Bible and the Quran. Timber Press, Inc. Portland, Oregon. Pp. 194–197.
- ↑ Weeks, A., et al. 2005. The phylogenetic history and biogeography of the frankincense and myrrh family (Burseraceae) based on nuclear and chloroplast sequence data. Molecular Phylogenetics and Evolution. 35:85–101.
- ↑ 8,0 8,1 8,2 8,3 8,4 Шаблон:Cite web
- ↑ Шаблон:Citation
- ↑ 10,0 10,1 Шаблон:Cite journal
- ↑ Шаблон:Cite web
- ↑ TRANS-MAD Development Parc National de Kirindy-Mite
- ↑ Шаблон:Cite news
- ↑ Шаблон:Cite web
- ↑ Шаблон:Cite web