Английская Википедия:Dudleya anomala
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Dudleya anomala is a rare species of succulent plant in the family Crassulaceae commonly known as the Todos Santos liveforever. With a dense, cushion-forming habit, this leaf succulent is characterized by elongated stems, slightly sticky leaves, and bell-shaped flowers with white, spreading petals. This species is native to Baja California, Mexico, and is found primarily on islands and one coastal locality.[1]
Description
Morphology
Dudleya anomala is a rosette-forming leaf succulent that grows in a caespitose habit, with the stems branching to form dense cushions of rosettes Шаблон:Convert in diameter. The stems are elongated and procumbent, and may reach up to Шаблон:Cvt long, and are usually Шаблон:Convert thick. Topping the stems are the rosettes, which contain 20 to 30 close-set leaves, that may become somewhat separated when the stem is rapidly elongating. The leaves are shaped strap-oblanceolate to strap-shaped, with acute tips. The lower surface of the leaf is convex, while the upper surface is only slightly convex. The leaf margins are obtuse or rounded, but not toward the base. The leaves measure Шаблон:Convert long by Шаблон:Convert wide, with the leaf bases Шаблон:Convert broad. The leaves are slightly viscid.[2]
The peduncle is Шаблон:Convert by Шаблон:Convert thick, and is covered in ascending bracts. The bracts are shaped narrowly deltoid-ovate to deltoid-lanceolate, with sharply acute tips. The lower bracts measure Шаблон:Convert long by Шаблон:Convert wide. The inflorescence is compact, with an overall rounded shape, and measures about Шаблон:Convert in diameter. The inflorescence usually has three major branches which may or may not subsequently rebranch once, and the terminal branches contain 3 to 5 flowers, held on pedicels Шаблон:Convert long.[2]
On the flower, the sepals are deltoid to deltoid-ovate with acute tips, measuring Шаблон:Convert long. The petals are somewhat spreading from the base, with the tips curving outward. Petals are narrowly ovate with acute tips, measuring Шаблон:Convert long by Шаблон:Convert wide, connate for Шаблон:Cvt or less. The petals are white, but somewhat flecked on the keel with red. The stamens measure about Шаблон:Convert long, and are adnate with the petals for Шаблон:Convert. The anthers are orange, and Шаблон:Cvt. The carpels are Шаблон:Convert and connate for about Шаблон:Cvt, spreading in age, with the styles Шаблон:Cvt long.[2]
Characteristics
D. anomala possesses slightly viscid (sticky) and odorous foliage, a trait only shared by the aptly-named Dudleya viscida.[3] In comparison, anomala has smaller leaves, thinner caudices, and smaller, simpler inflorescences.[2]
Taxonomy
Taxonomic history
Anstruther Davidson first described the species as Stylophyllum anomalum in 1928, based on a specimen collected by Robert Kessler from North Coronado Island and deposited in the herbarium of the Los Angeles County Museum by Davidson. Harald August Fröderström described Stylophyllum insulare based on a specimen collected by Marcus E. Jones from the Coronado Islands in 1926. This name later became a homonym, so it was changed to Stylophyllum coronatum. Philip A. Munz also observed this species on the Coronado Islands, referring to it as Echeveria virens.[2]
Reid Moran wrote a more complete description of the species in 1943, and recognized it as a Dudleya, creating the current combination.[2]
Phylogenetics
A 2013 phylogenetic analysis of D. anomala places it in a clade with Dudleya anthonyi.[4]
Distribution and habitat
Dudleya anomala is only found in the state of Baja California, Mexico and has an insular distribution on the Islas Coronados, Isla Todos Santos, and one location on the mainland, the extreme point of the Punta Banda. The range of D. anomala overlaps with those of Dudleya lanceolata, Dudleya candida, and Dudleya attenuata.[5][6][7]
Specimens of D. anomala observed by Reid Moran occurred on steep, north-facing cliffs on the Coronado Islands.[2] On the Isla Todos Santos, the plant is found densely covering rocks and cliffs on northern and eastern exposures.[8] On the Punta Banda, this species is found on steep, north-facing beach cliffs overlooking the Todos Santos Bay.[6] Populations of Dudleya anomala are consistently associated with Niebla ceruchoides.[9]
Gallery
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Cultivated plant with nascent inflorescences.
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At the University of California Botanical Garden, Berkeley, California
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At the Huntington Library.
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Detail of a flower and bud.
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Multiple flowers and buds.
References
External links
- ↑ Шаблон:Cite journal
- ↑ 2,0 2,1 2,2 2,3 2,4 2,5 2,6 Dudleya anomala (Davidson) Moran. Desert Plant Life. 14: 191. 1942.
- ↑ Шаблон:Cite eFloras
- ↑ Yost, J. M., Bontrager, M., McCabe, S. W., Burton, D., Simpson, M. G., Kay, K. M., & Ritter, M. (2013). Phylogenetic relationships and evolution in Dudleya (Crassulaceae). Systematic Botany, 38(4), 1096-1104.
- ↑ Oberbauer, T. A. (2002, March). Vegetation and flora of Islas Los Coronados, Baja California, México. In Proceedings of the Fifth California Islands Symposium (Vol. 29, pp. 212-223).
- ↑ 6,0 6,1 Mulroy, T. W., Rundel, P. W., & Bowler, P. A. (1979). The vascular flora of Punta Banda, Baja California Norte, Mexico. Madroño, 69-90.
- ↑ Uhl, C. H., & Moran, R. (1953). The cytotaxonomy of Dudleya and Hasseanthus. American Journal of Botany, 492-502.
- ↑ Шаблон:Cite journal
- ↑ Шаблон:Cite journal
