Английская Википедия:Elephantidae

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Elephantidae is a family of large, herbivorous proboscidean mammals collectively called elephants and mammoths. These are large terrestrial mammals with a snout modified into a trunk and teeth modified into tusks. Most genera and species in the family are extinct. Only two genera, Loxodonta (African elephants) and Elephas (Asian elephants), are living.

The family was first described by John Edward Gray in 1821,[1] and later assigned to taxonomic ranks within the order Proboscidea. Elephantidae has been revised by various authors to include or exclude other extinct proboscidean genera.

Description

Elephantids are distinguished from more primitive proboscideans like gomphotheres by their teeth, which have parallel lophs, formed from the merger of the cusps found in the teeth of more primitive proboscideans, which are bound by cement.[2] In later elephantids these lophs became narrow lamellae,[3] with the number of lophs/lamellae per tooth, as well as the tooth crown height (hypsodonty) increasing over time.[4] Elephantids chew using a proal jaw movement involving a forward stroke of the lower jaws, different from the oblique movement using side to side motion of the jaws in more primitive proboscideans.[5] The most primitive elephantid Stegotetrabelodon had a long lower jaw with lower tusks and retained permanent premolars similar to many gomphotheres, while modern elephantids lack permanent premolars, with the lower jaw being shortened (brevirostrine) and lower tusks being absent.[4]

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Classification

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Файл:Palaeoloxodon phylogeny.svg
Phylogeny of recent and Late Pleistocene elephantid species, including Palaeoloxodon and mammoths, showing the hybridisation between African forest elephants and Palaeoloxodon, after Palkopoulou et al. 2018
Файл:Comparative view of the human and elephant frame, Benjamin Waterhouse Hawkins, 1860.jpg
"Man, and the elephant" plate from Hawkins's A comparative view of the human and animal frame, 1860
Файл:Mammuthus meridionalis.JPG
Skeleton of Mammuthus meridionalis at the French Museum of Natural History

Some authors have suggested to classify the family into two subfamilies, Stegotetrabelodontinae, which is monotypic, only containing Stegotetrabelodon, and Elephantinae, containing all other elephantids.[4] Recent genetic research has indicated that Elephas and Mammuthus are more closely related to each other than to Loxodonta, with Palaeoloxodon closely related to Loxodonta. Palaeoloxodon also appears to have received extensive hybridisation with the African forest elephant, and to a lesser extent with mammoths.[6]

Extinct genera

Evolutionary history

Файл:ElephEvol.jpg
Evolution of elephants from the ancient Eocene (bottom) to the modern day (top)

Elephantids are thought to have evolved from gomphotheres, with some authors proposing the most likely ancestors to be African species of the "tetralophodont gomphothere" Tetralophodon.[7] The earliest members of the family, are known from the Late Miocene, around 9–10 million years ago.[8] The modern genera of elephants and mammoths had diverged from each other by the end of the Miocene. Elephantids began to migrate out of Africa during the Pliocene, with mammoths and Elephas arriving in Eurasia around 3–3.8 million years ago.[9] Around 1.5 million years ago, mammoths migrated into North America.[10] At the end of the Early Pleistocene, around 0.8 million years ago, Palaeoloxodon migrated out of Africa, becoming widespread across Eurasia, from Western Europe to Japan.[11] Palaeoloxodon and Mammuthus became extinct during the Late Pleistocene-Holocene, with the last population of mammoths persisting on Wrangel Island until around 4,000 years ago.[12]

See also

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References

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External links

Шаблон:Taxonbar Шаблон:Authority control

Шаблон:Proboscidea Genera

  1. Шаблон:Cite journal
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  4. 4,0 4,1 4,2 Шаблон:Citation
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  8. H. Saegusa, H. Nakaya, Y. Kunimatsu, M. Nakatsukasa, H. Tsujikawa, Y. Sawada, M. Saneyoshi, T. Sakai Earliest elephantid remains from the late Miocene locality, Nakali, Kenya Scientific Annals, School of Geology, Aristotle University of Thessaloniki, Greece VIth International Conference on Mammoths and Their Relatives, vol. 102, Grevena -Siatista, special volume (2014), p. 175
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